To determine if changes in the key bioclimatic parameters associated with rainfall seasonality can be quantified from fossil sequences and to distinguish them from changes in the total annual ...rainfall. Lake Masoko, southern Tanzania (9°20' S, 33°45' E, 840 m a.s.l.). Fossil pollen was extracted from a long and well-dated sedimentary sequence spanning the period 45,000-4400 cal. yr bp. The modern habitat, geographical distribution and climatic range (mean annual rainfall and temperature, and length of the dry season) of selected East African plants were used to infer past rainfall attributes for the most characteristic fossil pollen taxa identified from the sediments of Lake Masoko. In the Masoko pollen sequence it has been possible to identify changes in the length/severity of the dry season during the last 45,000 cal. yr bp, which are interpreted to reflect shifts in the mean position of the Intertropical Convergence Zone (ITCZ) over the southern tropics. We suggest that this bioclimatic parameter has been the main driver of the vegetation dynamics in this area. The major inferred change occurred at the Younger Dryas-Holocene transition c. 11,800 cal. yr bp, when semi-deciduous forest disappeared, being replaced in the lake catchment by woodland, which persists to the present day in spite of locally high rainfall. In tropical regions under the influence of the ITCZ, the position and strength of which determine the distribution of rainfall through the year, more attention must be paid to the impact of the length and intensity of the dry season on the modern and past distribution and dynamics of the vegetation. This climatic parameter is as important as the total annual amount of rainfall, and probably one of the most relevant in lowland areas.
Tropical climate is rapidly changing, but the effects of these changes on the geosphere are unknown, despite a likelihood of climatically-induced changes on weathering and erosion. The lack of long, ...continuous paleo-records prevents an examination of terrestrial responses to climate change with sufficient detail to answer questions about how systems behaved in the past and may alter in the future. We use high-resolution records of pollen, clay mineralogy, and particle size from a drill core from Lake Malawi, southeast Africa, to examine atmosphere-biosphere-geosphere interactions during the last deglaciation (∼ 18-9 ka), a period of dramatic temperature and hydrologic changes. The results demonstrate that climatic controls on Lake Malawi vegetation are critically important to weathering processes and erosion patterns during the deglaciation. At 18 ka, afromontane forests dominated but were progressively replaced by tropical seasonal forest, as summer rainfall increased. Despite indication of decreased rainfall, drought-intolerant forest persisted through the Younger Dryas (YD) resulting from a shorter dry season. Following the YD, an intensified summer monsoon and increased rainfall seasonality were coeval with forest decline and expansion of drought-tolerant miombo woodland. Clay minerals closely track the vegetation record, with high ratios of kaolinite to smectite (K/S) indicating heavy leaching when forest predominates, despite variable rainfall. In the early Holocene, when rainfall and temperature increased (effective moisture remained low), open woodlands expansion resulted in decreased K/S, suggesting a reduction in chemical weathering intensity. Terrigenous sediment mass accumulation rates also increased, suggesting critical linkages among open vegetation and erosion during intervals of enhanced summer rainfall. This study shows a strong, direct influence of vegetation composition on weathering intensity in the tropics. As climate change will likely impact this interplay between the biosphere and geosphere, tropical landscape change could lead to deleterious effects on soil and water quality in regions with little infrastructure for mitigation.
Celotno besedilo
Dostopno za:
DOBA, IZUM, KILJ, NUK, PILJ, PNG, SAZU, SIK, UILJ, UKNU, UL, UM, UPUK
To reconstruct the response of vegetation to abrupt climate changes during the last glacial we have compiled pollen records from the circum-Atlantic tropics between 23°N and 23°S from both marine and ...terrestrial sediment cores. Pollen data were grouped into mega-biomes to facilitate the comparison between the different records. Most tropical African records do not appear to register Dansgaard–Oeschger (D-O) variability, although there are vegetation changes during Heinrich Stadials (HS). There is a stronger signal of D-O and HS variability in the South American records. Records close to the modern northern and southern limits of the Intertropical Convergence Zone (ITCZ) show opposite trends in vegetation development during HS and D-O cycles. The pollen data from tropical South America corroborate the hypothesis of a southward shift in the migration pattern of the ITCZ and a reduction in the Atlantic Meridional Overturning Circulation (AMOC) during HS.
A 45,000‐year high‐resolution sedimentary record from Lake Masoko in Tanzania shows the climate in this part of East Africa to have been characterized by a short and less severe dry season during the ...Last Glacial Maximum. Moisture and lake‐level proxies, pollen assemblages and magnetic susceptibility, indicate that rainfall in the Masoko area was strongly controlled by low‐latitude insolation (i.e., precessional forcing). Such observations contrast with the climatic patterns previously reconstructed in East Africa for the last glacial. Indeed, widespread dry conditions are generally observed and are attributed to the predominant control of high latitude glacial forcing over insolation forcing on the tropical hydrology. However, during the Younger Dryas cold event, out of phase with regional insolation behaviour, wetter conditions prevailed in this area, suggesting that the rainfall belt over Africa was probably shifted to the South, bringing moisture to the southernmost tropics.
Impacts of global climate change on terrestrial ecosystems are imperfectly constrained by ecosystem models and direct observations. Pervasive ecosystem transformations occurred in response to warming ...and associated climatic changes during the last glacial-to-interglacial transition, which was comparable in magnitude to warming projected for the next century under high-emission scenarios. We reviewed 594 published paleoecological records to examine compositional and structural changes in terrestrial vegetation since the last glacial period and to project the magnitudes of ecosystem transformations under alternative future emission scenarios. Our results indicate that terrestrial ecosystems are highly sensitive to temperature change and suggest that, without major reductions in greenhouse gas emissions to the atmosphere, terrestrial ecosystems worldwide are at risk of major transformation, with accompanying disruption of ecosystem services and impacts on biodiversity.
Trees and shrubs in tropical Africa use the C-3 cycle as a carbon fixation pathway during photosynthesis, while grasses and sedges mostly use the C-4 cycle. Leaf-wax lipids from sedimentary archives ...such as the long-chain n-alkanes (e.g., n-C-27 to n-C-33) inherit carbon isotope ratios that are representative of the carbon fixation pathway. Therefore, n-alkane delta C-13 values are often used to reconstruct past C-3/C-4 composition of vegetation, assuming that the relative proportions of C-3 and C-4 leaf waxes reflect the relative proportions of C-3 and C-4 plants. We have compared the delta C-13 values of n-alkanes from modern C-3 and C-4 plants with previously published values from recent lake sediments and provide a framework for estimating the fractional contribution (areal-based) of C-3 vegetation cover (f(C3)) represented by these sedimentary archives. Samples were collected in Cameroon, across a latitudinal transect that accommodates a wide range of climate zones and vegetation types, as reflected in the progressive northward replacement of C-3-dominated rain forest by C-4-dominated savanna. The C-3 plants analysed were characterised by substantially higher abundances of n-C-29 alkanes and by substantially lower abundances of n-C-33 alkanes than the C-4 plants. Furthermore, the sedimentary delta C-13 values of n-C-29 and n-C-31 alkanes from recent lake sediments in Cameroon (-37.4%) to 26.5%) were generally within the range of delta C-13 values for C-3 plants, even when from sites where C-4 plants dominated the catchment vegetation. In such cases simple linear mixing models fail to accurately reconstruct the relative proportions of C-3 and C-4 vegetation cover when using the delta C-13 values of sedimentary n-alkanes, overestimating the proportion of C-3 vegetation, likely as a consequence of the differences in plant wax production, preservation, transport, and/or deposition between C-3 and C-4 plants. We therefore tested a set of non-linear binary mixing models using delta C-13 values from both C-3 and C-4 vegetation as end-members. The non-linear models included a sigmoid function (sine-squared) that describes small variations in the f(C3) values as the minimum and maximum delta C-13 values are approached, and a hyperbolic function that takes into account the differences between C-3 and C-4 plants discussed above. Model fitting and the estimation of uncertainties were completed using the Monte Carlo algorithm and can be improved by future data addition. Models that provided the best fit with the observed delta C-13 values of sedimentary n-alkanes were either hyperbolic functions or a combination of hyperbolic and sine-squared functions. Such non-linear models may be used to convert delta C-13 measurements on sedimentary n-alkanes directly into reconstructions of C-3 vegetation cover.
As part of a wider study of last glacial and deglacial climates in the Southern Hemisphere continents, we here review terrestrial and near-shore marine records from equatorial and southern Africa ...between 30,000 and 10,000
years ago (30–10
ka). This time interval covers the lead-up to the Last Glacial Maximum (LGM; 21
±
2
ka), the LGM proper, and the ensuing deglacial. Records selected for review needed to meet three requirements: continuity or near continuity over the period; a well-established chronology; and at least one but preferably several unambiguous proxy(ies).
We aim to show how regional climates of the sub-continent have responded to orbital forcing as opposed to other global glacial-interglacial boundary conditions, and how they are related to high latitude climates, sea and land surface conditions, positions of the Intertropical Convergence Zone (ITCZ) and of the westerly belt. Evidence of past climates derived from many independent proxies is given from west to southwest Africa (moisture from the Atlantic Ocean), then from equatorial East Africa to the southern subtropical summer rainfall domain (moisture mainly from the Indian Ocean). The LGM was cooler than today, and generally drier in the tropics. North of 8–9°S, glacial to Holocene increase in monsoonal precipitation, primarily related to orbitally-induced summer insolation in the northern hemisphere, occurred by steps of increasing amplitude (∼17–16, 14.5, 11.5
ka). Major wet–dry spells coincide with abrupt warm–cold events in high northern latitudes and related ITCZ migrations. In the southern tropics, the main post-glacial increase in tropical rainfall generally appears more gradual and in phase with Antarctic warming. Data suggest a restricted northward migration of the ITCZ and concentration of tropical rainfall well south of the Equator during the LGM and the Younger Dryas. Drier glacial conditions prevailed in southeastern Africa, while parts of southwestern Africa point to enhanced humidity during the LGM, suggesting that the winter westerly belt was either stronger than today or displaced further north possibly as a result of more extensive Antarctic sea-ice. Inferred African climatic fluctuations show the competing influences of tropical and high latitude climates of both hemispheres, and suggest changes in both meridional and zonal circulation modes. This review also reveals major geographical and methodological gaps, and a number of unresolved issues providing pointers for future research.
African ecosystems are at great risk. Despite their ecological and economic importance, long‐standing ideas about African forest ecology and biogeography, such as the timing of changes in forest ...extent and the importance of disturbance, have been unable to be tested due to a lack of sufficiently long records. Here, we present the longest continuous terrestrial record of late Quaternary vegetation from southern Africa collected to date from a drill core from Lake Malawi covering the last ~600,000 years. Pollen analysis permits us to investigate changes in vegetation structure and composition over multiple climatic transitions. We observe nine phases of forest expansion and collapse related to regional hydroclimate change. The development of desert, steppe and grassland vegetation during arid periods is likely dynamically linked to thresholds in regional hydrology associated with lake level and moisture recycling. Species composition of these dryland ecosystems varied greatly and is unlike the vegetation found at Malawi today, with assemblages suggesting strong Somali‐Masai affinities. Furthermore, nearly all semiarid assemblages contain low forest taxa abundances, suggesting that moist lowland gallery forests formed refugia along waterways during arid times. When the region was wet, forests were species‐rich and very high afromontane tree abundances suggest frequent widespread lowland colonization by modern high elevation trees. Furthermore, species composition varied little amongst forest phases until ~80 ka when disturbance tolerant tree taxa characteristic of the modern vegetation increased in abundance. The waxing and waning of forests has important implications for understanding the processes that control modern tropical vegetation biogeography as well as the environments of early humans across Africa. Finally, this work highlights the resilience of montane forests during previous warm intervals, which is relevant for future climate change; however, we point to a fundamental shift in disturbance regimes which are crucial for the structure and composition of modern East African landscapes.
Very little is known of African forest history to understand how forests may change in the future. This study presents 600,000 years of vegetation history from Lake Malawi showing massive reorganization of the vegetation due to changes in regional rainfall. We find that forest were widespread for much of the last 600,000 years, and that modern miombo woodland only emerged within the last 80 ka.
The Masoko crater-lake in southern Tanzania provides a continuous record of environmental changes covering the last 500 years. Multi-proxy studies were performed on a 52 cm sediment core retrieved ...from the deepest part of the lake. Magnetic, organic carbon, geochemical proxies and pollen assemblages indicate a dry climate during the ‘Little Ice Age’ (AD 1550–1850), confirming that the LIA in eastern Africa resulted in marked and synchronous hydrological changes. However, the direction of response varies between different African lakes (low versus high lake-levels), indicating strong regional contrasts that prevent the clear identification of climate trends over eastern Africa at this time. Inferred changes in Masoko lake-levels closely resemble the record of solar activity cycles, indicating a possible control of solar activity on the climate in this area. This observation supports previous results from East African lakes, and extends this relationship southward. Finally, anthropogenic impact is observed in the Masoko sediments during the last 60 years, suggesting that human disturbance significantly affected this remote basin during colonial and post-colonial times.
Diatom genera in many large East African lakes change little throughout the Holocene period suggesting relatively stable ecological conditions and some resilience to environmental change. Ecosystem ...stability is less common in smaller, more sensitive lakes, such as those within volcanic craters, where external impacts can cause abrupt and rapid fluctuations. A 4100-year diatom and cyanobacteria pigment record from Lake Massoko, a volcanic crater lake in southern Tanzania, is used to illustrate important switches in resource ratios following tephra deposition 1190 years ago. It is hypothesized that the tephra reduced the rate of P diffusion from the sediments and increased the Si:P ratio in the lake. A period of acute change in planktonic diatom communities resulted from the tephra impact and lasted c. 110 years. The magnitude of the change shown by the diatoms and their slow recovery from the tephra may be due in part to a coincident fall in lake level caused by a reduction in regional rainfall. The statistical significance of the tephra impact relative to that of catchment and climate change has been tested using variance partitioning and rate-of-change analysis. Multiproxy indicators show an important period of positive water balance 1700 ago and a relatively dry episode persisting between 1000 and 400 years ago. The lake ecosystem is shown to be highly sensitive to both climate change and tephra deposition.
Celotno besedilo
Dostopno za:
DOBA, IZUM, KILJ, NUK, OILJ, PILJ, PNG, SAZU, UILJ, UKNU, UL, UM, UPUK
PDF
