Mechanisms of selection for domestication traits in cereals and other annual plants are commonly explained from agro-technological and genetic perspectives. Since archaeobotanical data showed that ...domestication processes were slow and protracted, research focused on genetic constraints and hypothetical ‘non-selective’ management regimes to explain the low selection rates. I argue that these factors only partially explain the observed patterns and develop a model that contextualises the archaeobotanical data in their socio-economic settings. I propose that developments towards individual storage by small household units and the gradual increase in storage capacities with the development of extended households represent key factors for establishing the conditions for selection, as these practices isolated individually managed and stored cereal subpopulations and gradually reduced the need to replenish grain stocks with grains from unmanaged populations. This genetic isolation resulted in stronger and more persistent selection rates and facilitated the genetic fixation of domestication traits on a population level. Moreover, individual storage facilities within buildings reflect gradual developments towards households as the social units that mobilised agricultural labour, which negotiated new sharing principles over cultivated resources and drove the intensification of cultivation practices. In this sense, selection rates and the slow domestication process can be understood as a function of limited food sharing networks and increased labour-inputs into early arable environments—socio-economic processes that also unfolded gradually over a protracted period of time.
The present study investigates the occurrence of wild grasses at Epipalaeolithic and aceramic Neolithic sites in the Near East in order to assess their role in subsistence economies alongside the ...emergence of cereal cultivation. We use Chogha Golan in the foothills of the central Zagros Mountains (ca. 11.7-9.6 ka cal. BP) as a case study, where the archaeobotanical data suggest the frequent exploitation of a complex of wild grasses for almost 2,000 years. Domesticated emmer replaced these wild grasses as the major food resources towards the end of occupation at the site (ca. 9.8 ka cal. BP). We discuss possible implications of this development and conclude that the traditional concept of pre-domestication cultivation seems unsuited for explaining the patterns from Chogha Golan. These data are in good accordance with the overall picture in the Zagros Mountains, where wild grasses were routinely gathered throughout the early Holocene. In contrast, wild grasses were gradually replaced by wild cereals in the Levantine corridor since the end of the Pleistocene. However, several sites located in this region provide evidence for a continuous exploitation of wild grasses alongside emerging cereal cultivation and most of these taxa were part of the earliest segetal floras that evolved with the appearance of domestic cereals throughout the 11th millennium cal. BP. Some sites contemporary to the Pre-Pottery Neolithic B still provide evidence for the usage of wild grasses, which possibly reflects the utilization of edible arable weeds and continuous gathering of wild grasses by more mobile groups.
Celotno besedilo
Dostopno za:
DOBA, IZUM, KILJ, NUK, PILJ, PNG, SAZU, SIK, UILJ, UKNU, UL, UM, UPUK
Reconstructing the domestication process of cereals is a basic requirement for the understanding of the Early Neolithic in the Near East and how agriculture emerged. Although there is general ...agreement on the criteria of differentiating wild from domesticated cereals, their application to material from aceramic Neolithic sites is problematic for reasons of preservation and diversity of early transitional chaff remains. In the present study we established an identification key for the distinction of wild and domestic emmer wheat (Triticum turgidum subsp. dicoccoides/dicoccum). General morphological analyses as well as experimental charring and measurements on wild and domestic emmer from the Fertile Crescent were conducted to track the main features that distinguish the two forms. Wild emmer can be differentiated from domestic emmer using longitudinal sections through rachises. The scar morphology of wild emmer specimens with a rough upper abscission scar is distinct from domestic emmer. In addition, two of the measuring tracks distinguish between domestic emmer and its progenitor. These results were applied to archaeological specimens from the aceramic Neolithic site of Chogha Golan (Ilam Province, Iran), which was excavated in 2009 and 2010 by a team of the Tübingen Iranian Stone Age Research Project. The carbonized emmer remains dating to about 9800BP comprise domestic-type and wild-type rachises, which is typical for cereal assemblages from aceramic Neolithic deposits.
•Morphology of wild and domestic emmer wheat accessions was analyzed.•Both are distinguishable using the upper abscission zone in longitudinal section.•Rough scars among wild emmer are distinct from rough scars of domestic emmer.•Domestic-type emmer dating to about 9800B.P. was identified at Chogha Golan.
Reconstructing the origins of plant cultivation in southwest Asia is crucial for understanding associated processes such as the emergence of sedentary communities and domesticated crops. Among the ...criteria archaeobotanists developed for identifying the earliest plant cultivation, the presence of potential arable weeds found in association with wild cereal and legume remains has been used as a basis for supporting models of prolonged wild plant cultivation before domesticated crops appear. However, the proposed weed floras mainly consist of genus-level identifications that do not differentiate between arable weeds and related species that characterise non-arable habitats. Here we test, for the first time, whether the potential arable weed taxa widely used to identify wild plant cultivation also occur in non-cultivated wild cereal populations. Based on modern survey data from the southern Levant we show that the proposed weed taxa characterise both grasslands and fields. Our findings, therefore, do not support the use of these taxa for reconstructing early cultivation. Instead, for future studies we suggest an approach based on the analysis of plant functional traits related to major agroecological variables such as fertility and disturbance, which has the potential to overcome some of the methodological problems.
Process philosophy offers a metaphysical foundation for domestication studies. This grounding is especially important given the European colonialist origin of 'domestication' as a term and 19th ...century cultural project. We explore the potential of process archaeology for deep-time investigation of domestication relationships, drawing attention to the variable pace of domestication as an ongoing process within and across taxa; the nature of domestication 'syndromes' and 'pathways' as general hypotheses about process; the importance of cooperation as well as competition among humans and other organisms; the significance of non-human agency; and the ubiquity of hybrid communities that resist the simple wild/domestic dichotomy.
Celotno besedilo
Dostopno za:
DOBA, IZUM, KILJ, NUK, PILJ, PNG, SAZU, SIK, UILJ, UKNU, UL, UM, UPUK
The protracted domestication model posits that wild cereals in southwest Asia were cultivated over millennia before the appearance of domesticated cereals in the archaeological record. These ...'pre-domestication cultivation' activities are widely understood as entailing annual cycles of soil tillage and sowing and are expected to select for domestic traits such as non-shattering ears. However, the reconstruction of these practices is mostly based on indirect evidence and speculation, raising the question of whether pre-domestication cultivation created arable environments that would select for domestic traits. We developed a novel functional ecological model that distinguishes arable fields from wild cereal habitats in the Levant using plant functional traits related to mechanical soil disturbance. Our results show that exploitation practices at key pre-domestication cultivation sites maintained soil disturbance conditions similar to untilled wild cereal habitats. This implies that pre-domestication cultivation did not create arable environments through regular tillage but entailed low-input exploitation practices oriented on the ecological strategies of the competitive large-seeded grasses themselves.
In this paper we discuss the plant-based subsistence economy during the formation of archaeological horizons (AH) II and I at the aceramic Neolithic site of Chogha Golan, Iran. The deposits date to ...between 9,800 and 9,600 cal bp. In order to reconstruct subsistence practices and their development reliably, we conducted a taphonomic analysis to identify factors that influenced the composition of the archaeobotanical assemblage. The flotation samples derive from two excavation areas in the centre of the tell, the deep sounding and area A. Using correspondence analysis, we link the biased composition of the plant remains from AH I to their relatively poor preservation. Two different sampling strategies applied in excavation area A also affected the composition of the samples. In contrast, we did not find compositional differences among the samples from AH II of both excavation areas. Our results emphasize the need for taphonomic analyses prior to interpreting the taxonomic composition of charred archaeobotanical assemblages. Considering these results, we discuss the subsistence economy of Chogha Golan. Domestic emmer wheat was cultivated from AH II onwards. Wild barley, Aegilops sp., lentils, peas and various vetches may have been cultivated as well. This spectrum of typical Neolithic food plants was supplemented by a high diversity of other potential wild food resources, including medium and small-seeded grasses, Pistacia, Bolboschoenus glaucus, Malva and Brassicaceae. A compilation of ethnobotanical data, mainly from the Near and Middle East, represents the basis for assessing the potential uses of the wild plants.
The so-called Triticoid-type grains are known from several prehistoric sites in southwest Asia and their identification has long been unclear. They resemble the grains of wheats and researchers ...suggested they may represent an extinct Triticeae species, possibly closely related to wild crop progenitors. In this study we identify the Triticoid-type grains as
Heteranthelium piliferum
(Banks & Sol.) Hochst. and describe the key identification criteria. The identification is based on morphological analyses of modern and archaeological material from several grass species and was first achieved with well-preserved specimens from Early Neolithic Chogha Golan, Iran. We further examined the Triticoid-type grains from recently excavated samples from Early Neolithic Ganj Dareh, Iran, and archived samples from Late Chalcolithic and Late Bronze Age Tell Brak in northeast Syria, confirming their identification as
H. piliferum
. Based on the study of herbarium specimens at Royal Botanic Gardens Kew, London, we provide a detailed distribution map and review the species’ biology and ecological adaptations. Collected and cultivated herbarium specimens were analysed in order to understand the high phenotypic plasticity of the growth habit, its correlation with environmental variables and its relation to grain size. In order to understand the high morphological variability of the charred Triticoid-type grains from archaeological deposits, we assessed the effects of experimental carbonisation at different temperatures on grains of
H. piliferum
,
Triticum dicoccum
,
T. thaoudar
and
Secale vavilovii
. In light of the present study, we discuss the relevance of
H. piliferum
for reconstructing prehistoric subsistence strategies.
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