The role of Iran as a center of origin for domesticated cereals has long been debated. High stratigraphie resolution and rich archaeological remains at the aceramic Neolithic site of Chogha Golan ...(Ham Province, present-day Iran) reveal a sequence ranging over 2200 years of cultivation of wild plants and the first appearance of domesticated-type species. The botanical record from Chogha Golan documents how the inhabitants of the site cultivated wild barley (Hordeum spontaneum) and other wild progenitor species of modern crops, such as wild lentil and pea. Wild wheat species (Triticum spp.) are initially present at less than 10% of total plant species but increase to more than 20% during the last 300 years of the sequence. Around 9800 calendar years before the present, domesticated-type emmer appears. The archaeobotanical remains from Chogha Golan represent the earliest record of long-term plant management in Iran.
The present study investigates the occurrence of wild grasses at Epipalaeolithic and aceramic Neolithic sites in the Near East in order to assess their role in subsistence economies alongside the ...emergence of cereal cultivation. We use Chogha Golan in the foothills of the central Zagros Mountains (ca. 11.7-9.6 ka cal. BP) as a case study, where the archaeobotanical data suggest the frequent exploitation of a complex of wild grasses for almost 2,000 years. Domesticated emmer replaced these wild grasses as the major food resources towards the end of occupation at the site (ca. 9.8 ka cal. BP). We discuss possible implications of this development and conclude that the traditional concept of pre-domestication cultivation seems unsuited for explaining the patterns from Chogha Golan. These data are in good accordance with the overall picture in the Zagros Mountains, where wild grasses were routinely gathered throughout the early Holocene. In contrast, wild grasses were gradually replaced by wild cereals in the Levantine corridor since the end of the Pleistocene. However, several sites located in this region provide evidence for a continuous exploitation of wild grasses alongside emerging cereal cultivation and most of these taxa were part of the earliest segetal floras that evolved with the appearance of domestic cereals throughout the 11th millennium cal. BP. Some sites contemporary to the Pre-Pottery Neolithic B still provide evidence for the usage of wild grasses, which possibly reflects the utilization of edible arable weeds and continuous gathering of wild grasses by more mobile groups.
Celotno besedilo
Dostopno za:
DOBA, IZUM, KILJ, NUK, PILJ, PNG, SAZU, SIK, UILJ, UKNU, UL, UM, UPUK
Abstract
Although Middle Paleolithic (MP) hominin diets consisted mainly of ungulates, increasing evidence demonstrates that hominins at least occasionally consumed tortoises, birds, leporids, fish, ...and carnivores. Until now, the MP zooarchaeological record in the Zagros Mountains has been almost exclusively restricted to ungulates. The narrow range of hominin prey may reflect socioeconomic decisions and/or environmental constraints, but could also result from a research bias favoring the study of large prey, since archaeologists have undertaken no systematic taphonomic analyses of small game or carnivores in the region. Here, we report on the first comprehensive taphonomic analysis of an MP faunal assemblage from Ghar-e Boof (∼ 81–45 kyr), a Late Pleistocene site in the southern Zagros of Iran. Anthropogenic bone surface modifications point to hominins as the main agent of accumulation. Hominins preyed primarily on ungulates, particularly wild goat. However, we also found evidence for MP hominin exploitation of carnivores and tortoises at the site. Although small game represents only a minor portion of the diet, our results suggest that the hunting behavior of MP hominins in the Zagros was more diverse than previously thought, similar to what we find elsewhere in Eurasia.
The so-called Triticoid-type grains are known from several prehistoric sites in southwest Asia and their identification has long been unclear. They resemble the grains of wheats and researchers ...suggested they may represent an extinct Triticeae species, possibly closely related to wild crop progenitors. In this study we identify the Triticoid-type grains as
Heteranthelium piliferum
(Banks & Sol.) Hochst. and describe the key identification criteria. The identification is based on morphological analyses of modern and archaeological material from several grass species and was first achieved with well-preserved specimens from Early Neolithic Chogha Golan, Iran. We further examined the Triticoid-type grains from recently excavated samples from Early Neolithic Ganj Dareh, Iran, and archived samples from Late Chalcolithic and Late Bronze Age Tell Brak in northeast Syria, confirming their identification as
H. piliferum
. Based on the study of herbarium specimens at Royal Botanic Gardens Kew, London, we provide a detailed distribution map and review the species’ biology and ecological adaptations. Collected and cultivated herbarium specimens were analysed in order to understand the high phenotypic plasticity of the growth habit, its correlation with environmental variables and its relation to grain size. In order to understand the high morphological variability of the charred Triticoid-type grains from archaeological deposits, we assessed the effects of experimental carbonisation at different temperatures on grains of
H. piliferum
,
Triticum dicoccum
,
T. thaoudar
and
Secale vavilovii
. In light of the present study, we discuss the relevance of
H. piliferum
for reconstructing prehistoric subsistence strategies.
Reconstructing the domestication process of cereals is a basic requirement for the understanding of the Early Neolithic in the Near East and how agriculture emerged. Although there is general ...agreement on the criteria of differentiating wild from domesticated cereals, their application to material from aceramic Neolithic sites is problematic for reasons of preservation and diversity of early transitional chaff remains. In the present study we established an identification key for the distinction of wild and domestic emmer wheat (Triticum turgidum subsp. dicoccoides/dicoccum). General morphological analyses as well as experimental charring and measurements on wild and domestic emmer from the Fertile Crescent were conducted to track the main features that distinguish the two forms. Wild emmer can be differentiated from domestic emmer using longitudinal sections through rachises. The scar morphology of wild emmer specimens with a rough upper abscission scar is distinct from domestic emmer. In addition, two of the measuring tracks distinguish between domestic emmer and its progenitor. These results were applied to archaeological specimens from the aceramic Neolithic site of Chogha Golan (Ilam Province, Iran), which was excavated in 2009 and 2010 by a team of the Tübingen Iranian Stone Age Research Project. The carbonized emmer remains dating to about 9800BP comprise domestic-type and wild-type rachises, which is typical for cereal assemblages from aceramic Neolithic deposits.
•Morphology of wild and domestic emmer wheat accessions was analyzed.•Both are distinguishable using the upper abscission zone in longitudinal section.•Rough scars among wild emmer are distinct from rough scars of domestic emmer.•Domestic-type emmer dating to about 9800B.P. was identified at Chogha Golan.
The beginnings of agriculture throughout the Fertile Crescent are still not completely understood, particularly at the eastern end of the Fertile Crescent in the area of modern Iran. Archaeobotanical ...samples from Epipalaeolithic/PPNA Körtik Tepe in southeastern Turkey and from the Pre-Pottery Neolithic sites of Chogha Golan and East Chia Sabz in south western Iran were studied in order to define the status of cultivation at these sites. Preliminary results show the presence of abundant wild progenitor species of crops at the Iranian sites before 10600 cal. B.P.: , and very few wild progenitor species at Körtik Tepe dated to 11700–11250 cal. B.P.: The Iranian sites also indicate size increase of wild barley grain across a sequence of 400 years through either cultivation or changing moisture conditions.
Ghār-e Boof is a Paleolithic cave site in Iran well known for its rich early Upper Paleolithic Rostamian assemblages. The site is located on the edge of the Dasht-e Rostam plain in the southern ...Zagros. Recent excavations by the members of the Tübingen-Iranian Stone Age Research Project at Ghār-e Boof also recovered well-stratified Middle Paleolithic assemblages. Here, we provide the first detailed luminescence chronology for the Middle and Upper Paleolithic of the site. More generally, our work is the first luminescence chronology for a Middle and Upper Paleolithic site in the Zagros Mountains region in Iran. The luminescence ages for the Upper Paleolithic of Ghār-e Boof agree with published 14C dates. We applied Bayesian models specifically designed for luminescence dating using the R package ‘BayLum’ to incorporate the well-established stratigraphic constraints, as well as the published 14C ages with our optically stimulated luminescence (OSL) ages to improve the precision of the chronological framework. The Bayesian chronology shows a significantly improved precision of the OSL ages in particular for the upper part of the sequence where 14C ages were available. The Bayesian OSL ages for the Rostamian horizons, archaeologicalhorizon (AH) III (a-b-c), and AH IV, fall in the range of 37–42 ka (68% credible interval CI). Moreover, we determined a series of dates between 45 and 81 ka (68% CI) for the Middle Paleolithic strata from AH IVd to AH VI. Our results point to a demographic shift in the populations responsible for the Middle Paleolithic and the Rostamian within three millennia. This major technological change accompanied by the rise of symbolic artifacts such as personal ornaments, may or may not reflect a replacement of Neanderthals by modern humans. While we are confident that the Rostamian was made by modern humans, available information does not allow us to be sure who made the local Middle Paleolithic.
Ghar-e Boof is a Paleolithic cave site in Iran, known for being the type locality for the early Upper Paleolithic (UP) Rostamian lithic industries. Defining the Rostamian cultural group has ...significant archaeological implications for the Zagros Mountains. First, it highlights the cultural diversity of the UP in the Zagros. Second, it evinces a more complex scenario for the spread of anatomically modern humans (AMHs) across Southwest Asia. Despite this, little is known about human subsistence strategies during the Rostamian. In this paper, we present the results of a zooarchaeological analysis that allows us to reconstruct prey choice, transport decisions, and carcass processing at Ghar-e Boof. Wild sheep/goat was the main prey at the site, but other taxa such as partridges, tortoises, and gazelles represented important dietary supplements. Mortality data indicate that Rostamian hunters primarily targeted prime-aged adult ungulates. Anatomical representation also shows that there was no selective transport of animal carcasses to the site, so hunting activities most likely took place nearby. Although many specimens in the assemblage were covered by thick concretions, the presence of cut marks demonstrates defleshing, filleting, dismemberment, and even the removal of internal organs of prey. Percussion impacts and cone fractures also suggest the processing of bones for marrow. Chronologically, moving from the oldest to the youngest layers, there is a decrease in large game in parallel with a progressive increase in small, fast-moving animals. Following the prey choice model of optimal foraging theory, this diachronic variation might track regional resource stress or an increase in hunting pressure. Our results, along with the analysis of lithic artifacts and changes in the rates of sedimentation during the Rostamian sequence, indicate an increase in occupation intensity through time. Thus, the archaeological record of Ghar-e Boof and the ubiquity of UP sites across the Dasht-e Rostam region most probably reflect population growth within a few millennia after the arrival of AMHs.
•First zooarchaeological study of the Rostamian sequence of Ghar-e Boof (42–35 ka cal BP).•Human exploitation of diverse animal prey.•Preferential hunting of prime-aged caprines near the site.•Evidence for changes in hunting pressure and site occupation intensity.
The excavation of Ghar-e Boof, a cave in the Zagros Mountains, places the site not only at the center of discussion on the transition to and development of its regional lithic tradition: the ...Rostamian (37,000–31,000 BP), but also in differentiating between plants used by humans or mere traces of the surrounding vegetation. The large pulses of
Lathyrus
or
Vicia
sp. recovered from this shallow cave in the southwest of Iran may, for instance, represent food collected from wild stands already in the early Upper Palaeolithic. The seeds of barley (
Hordeum
sp.), although not all clearly domesticated, are without doubt signs of disturbance or bioturbation since the historic era. Analysis of cave deposits over 30,000 years old raise a number of methodological and interpretive challenges. Human, taphonomic, or biomechanical disturbances impact the deposition of plant remains, as well as affect the composition of the assemblages, undermining spatial and ecological examination of the data set. Comprehension of provenance of the samples, site genesis, and matrix development, through detailed micromorphological and stratigraphical studies, is thus suggested in conjunction with the archaeobotanical analyses, to identify disturbances, define their cause, and treat them appropriately. Numerical studies and ecological interpretations of climate, vegetation composition, or indications of human activity therefore follow specific criteria discussed here. Despite signs of disturbance in the plant assemblages, archaeobotanic research can lead to recognition of environmental conditions, plausible human subsistence, site use and seasonality, and sound vegetation description.
The Zagros Mountains represent a strategic geographic region near the junction of Africa, Europe, and Asia, and valleys within the mountain range likely correspond to migration corridors used by ...archaic and modern humans. Ghar-e Boof, a key archeological site in the southern Zagros, is located at an altitude of 905 m.a.s.l. in the Dasht-e Rostam region of the Fars Province, Iran. On the basis of lithic artifacts and radiocarbon and luminescence dating, the Late Pleistocene archeological record of the site spans from the Middle Paleolithic (MP) to the Late Epipaleolithic. Ghar-e Boof includes find horizons of the early Upper Paleolithic (UP) Rostamian dating to 42–35 ka cal. BP. These Rostamian-UP assemblages are typically attributed to modern humans. The high-resolution stratigraphy and microvertebrate record offer a unique opportunity for reconstructing the paleoenvironment of the southern part of the Zagros Mountains during the Late Pleistocene and contribute to a better understanding of long-term hominin settlement dynamics and cultural adaptations in this area. The current paper presents the first analysis of the microvertebrate remains recovered by the Tübingen-Iranian Stone Age Research Project (TISARP) team. All groups of small vertebrates, including fish, amphibians, squamate reptiles, birds, and micromammals, are present in the assemblage from Ghar-e Boof (NISP = 755). The key micromammal taxa used to reconstruct the paleoenvironment during the MP and UP include Libyan jird (
Meriones
cf.
libycus
), Vinogradov’s jird (
Meriones vinogradovi
), Indian gerbil (
Tatera indica
), Brandt’s hedgehog (
Paraechinus
cf
. hypomelas
), house mouse
(Mus musculus
), and mole vole (
Ellobius
sp.). Other taxa, such as amphibians (toad,
Bufo
/
Bufotes
sp.), fish (nase,
Chondrostoma
sp.), and squamate reptiles (agamids, lizards, Eastern Montpelier snake—
Malpolon insignitus
—and viper), also support the presence of an environment mainly characterized by warm, arid conditions with open, dry meadows, rocky terrain, and water sources close to the site throughout most of the sequence. During the shift from MP to UP, we record Afghan pika (
Ochotona
cf.
rufescens
), while all rodents, except very few remains of the Persian vole (
Microtus
cf.
irani
) and the Libyan jird (
M.
cf.
lybicus
) and some non-hibernating vertebrate species, are absent and reappear in later UP horizons. The presence of pikas could reflect a period of slightly colder and/or drier conditions. While the record from Ghar-e Boof is the only dataset from the southern Zagros, previous studies on microvertebrates from the central and northern Zagros show no clear change during the shift from the MP to the UP. Data from sediment cores and pollen analyses document numerous environmental and climatic fluctuations during the Late Pleistocene of the Zagros. The microvertebrate record from Ghar-e Boof improves our understanding of paleoenvironments during the Late Pleistocene and opens new opportunities to reconstruct the ecological conditions that helped to shape hominin settlement dynamics in the region.
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