Social-ecological fit has been a popular approach to assessing the connectedness between social and ecological systems in environmental governance. However, the role of projects in social-ecological fit has yet to be explicitly problematized and conceptualized. Given the centrality of projects - i.e. temporally limited organizations that bring various actors into collaboration to pursue clearly defined goals and tasks - in environmental governance, this is a serious shortcoming in the literature. In this paper, we fill this gap by drawing on the hitherto unconnected literatures on projectified environmental governance and social-ecological fit. We then assess the extent to which fit can be achieved in projectified environmental governance. To do this, we develop a novel framework for assessing the vertical and horizontal dimensions of social, spatial, and temporal fit and conduct an empirical study of the European Union’s LIFE Program and environmental projects funded by the program in Estonia. Our results suggest that the spatial and temporal fit of projects is contingent on social fit, i.e., the alignment of interests and needs of project-related actors. Frictions between various levels of decision-making also condition the possibilities of achieving fit horizontally across stakeholder groups and ecological systems as well as of sustaining project results over time. •Socio-ecological fit is a widely debated topic in environmental governance.•A critical analysis of the role of projects in social-ecological fit is currently lacking.•We explore social-ecological fit of Estonian EU LIFE projects.•The interconnectedness between different forms of fit needs to be addressed.•Pursuing socio-ecological fit is a multi-scale and multi-level effort.