Mitochondria play central roles in cellular energetics, metabolism and signalling. Efficient respiration, mitochondrial quality control, apoptosis and inheritance of mitochondrial DNA depend on the ...proper architecture of the mitochondrial membranes and a dynamic remodelling of inner membrane cristae. Defects in mitochondrial architecture can result in severe human diseases affecting predominantly the nervous system and the heart. Inner membrane morphology is generated and maintained in particular by the mitochondrial contact site and cristae organizing system (MICOS), the F1Fo‐ATP synthase, the fusion protein OPA1/Mgm1 and the nonbilayer‐forming phospholipids cardiolipin and phosphatidylethanolamine. These protein complexes and phospholipids are embedded in a network of functional interactions. They communicate with each other and additional factors, enabling them to balance different aspects of cristae biogenesis and to dynamically remodel the inner mitochondrial membrane. Genetic alterations disturbing these membrane‐shaping factors can lead to human pathologies including fatal encephalopathy, dominant optic atrophy, Leigh syndrome, Parkinson's disease and Barth syndrome.
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We studied the properties of the neutral gas in a sample of 38 local luminous and ultra luminous infrared galaxies (ULIRGs, 51 individual galaxies at z ≤ 0.09), which mainly covers the less explored ...LIRG luminosity range. This study is based on the analysis of the spatially integrated and spatially resolved spectra of the NaDλλ 5890, 5896 Å feature obtained with the integral field unit (IFU) of VIMOS at the Very Large Telescope. Analyzing spatially integrated spectra, we find that the contribution of the stars to the observed NaD equivalent width is small (<35%) for about half of the sample, and therefore this feature is dominated by inter stellar medium (ISM) absorption. After subtracting the stellar contribution, we find that the pure-ISM integrated spectra generally show blueshifted NaD profiles, indicating neutral gas outflow velocities, V, in the range 65−260 km s-1. Excluding the galaxies with powerful AGNs, V shows a dependency with the star formation rate (SFR) of the type V ∝ SFR0.15, which is in rather good agreement with previous results. The spatially resolved analysis could be performed for 40 galaxies, 22 of which have neutral gas velocity fields dominated by noncircular motions with signatures of cone-like winds. However, a large number of targets (11/40) show disk rotation signatures. Based on a simple model, we found that the wind masses are in the range 0.4−7.5 × 108 M⊙, reaching up to ~3% of the dynamical mass of the host. The mass rates are typically only ~0.2−0.4 times the corresponding global SFR indicating that, in general, the mass loss is too small to slow down the star formation significantly. In the majority of cases, the velocity of the outflowing gas is not sufficient to escape the host potential well and, therefore, most of the gas rains back into the galaxy disk. On average V/vesc is higher in less massive galaxies, confirming that the galaxy mass has a primary role in shaping the recycling of gas and metals. The comparison between the wind power and kinetic power of the starburst associated with SNe indicates that only the starburst could drive the outflows in nearly all the ULIRGs galaxies, as the wind power is generally lower than 20% of the kinetic power supplied by the starburst. The contribution of an active galactic nuclei (AGN) is, in principle, significant in two cases.
We study the kinematic properties of the ionised gas outflows and ambient interstellar medium (ISM) in a large and representative sample of local luminous and ultra-luminous infrared galaxies ...(U/LIRGs) (58 systems, 75 galaxies) at galactic and sub-galactic (i.e., star-forming clumps) scales, thanks to integral field spectroscopy (IFS)-based high signal-to-noise integrated spectra. The velocity dispersion of the ionized ISM in U/LIRGs (⟨ σ ⟩ ~ 70 km s-1) is larger than in lower luminosity local star-forming galaxies (⟨ σ ⟩ ~ 25 km s-1). While for isolated disc LIRGs star formation appears to sustain turbulence, gravitational energy release associated with interactions and mergers plays an important role in driving σ in the U/LIRG range. We find that σ has a dependency on the star formation rate density (ΣSFR), which is weaker than expected if it were driven by the energy released by the starburst. The relatively small role of star formation (SF) driving the σ in U/LIRGs is reinforced by the lack of an increase in σ associated with high luminosity SF clumps. We also find that the impact of an active galactic nucleus (AGN) in ULIRGs is strong, increasing on average σ by a factor 1.5. Low-z U/LIRGs cover a range of velocity dispersion (σ ~ 30 to 100 km s-1) and star formation rate density (ΣSFR ~ 0.1 to 20 M⊙ yr-1 kpc-2) similar to those of high-z SFGs. Moreover, the observed weak dependency of σ on ΣSFR for local U/LIRGs (σ ∝ ΣSFR+0.06) is in very good agreement with that measured in some high-z samples. The presence of ionized gas outflows in U/LIRGs seems universal based on the detection of a broad, usually blueshifted, Hα line. The observed dependency of the maximum velocity of the outflow (Vmax) on the star formation rate (SFR) is of the type Vmax(non − AGN) ∝ SFR(LIR)+ 0.24. We find that AGNs in U/LIRGs are able to generate faster (~×2) and more massive (~× 1.4) ionized gas outflows than pure starbursts. The derived ionized mass loading factors (η) are in general below 1, with only a few AGNs above this limit. The escaping gas fraction is low with only less massive (log(Mdyn/M⊙) < 10.4) U/LIRGs having outflowing terminal velocities higher than their escape velocities, and more massive galaxies retaining the gas, even if they host an AGN. The observed average outflow properties in U/LIRGs are similar to high-z galaxies of comparable SFR. However, while high-z galaxies seem to require ΣSFR > 1 M⊙ yr-1 kpc-2 for launching strong outflows, this threshold is not observed in low-z U/LIRGs even after correcting for the differential fraction of the gas content. In the bright SF clumps found in LIRGs, ionized gas outflows appear to be very common (detection rate over 80%). Their observed properties are less extreme than those associated with the entire galaxy. The clumps in LIRGs follow the general size-L-σ scaling relations found for low- and high-z clumps, though they are in general smaller, less luminous, and are characterized by lower σ than at high-z. For a given observed (no internal extinction correction applied) star formation surface density, outflows in LIRG clumps would be about one to two orders of magnitude less energetic than the outflows launched by clumps in high-z SF galaxies.
Aims. We investigate the fueling and the feedback of nuclear activity in the nearby (D = 14 Mpc) Seyfert 2 barred galaxy NGC 1068 by studying the distribution and kinematics of molecular gas in the ...torus and its connections to the host galaxy disk. Methods. We used the Atacama Large Millimeter Array (ALMA ) to image the emission of a set of molecular gas tracers in the circumnuclear disk (CND) and the torus of the galaxy using the CO(2–1), CO(3–2), and HCO+(4–3) lines and their underlying continuum emission with high spatial resolutions (0.03″ − 0.09″ ≃ 2 − 6 pc). These transitions, which span a wide range of physical conditions of molecular gas (n(H2)⊂103 − 107 cm−3), are instrumental in revealing the density radial stratification and the complex kinematics of the gas in the torus and its surroundings. Results. The ALMA images resolve the CND as an asymmetric ringed disk of D ≃ 400 pc in size and ≃1.4 × 108 M⊙ in mass. The CND shows a marked deficit of molecular gas in its central ≃130 pc region. The inner edge of the ring is associated with the presence of edge-brightened arcs of NIR polarized emission, which are identified with the current working surface of the ionized wind of the active galactic nucleus (AGN). ALMA proves the existence of an elongated molecular disk/torus in NGC 1068 of Mtorusgas ≃ 3 × 105 M⊙ M torus gas ≃ 3 × 10 5 M ⊙ $ M_{\mathrm{torus}}^{\mathrm{gas}}\simeq3\times10^{5}\,M_{{\odot}} $ , which extends over a large range of spatial scales D ≃ 10 − 30 pc around the central engine. The new observations evidence the density radial stratification of the torus: the HCO+(4–3) torus, with a full size DHCO+(4 − 3) = 11 ± 0.6 pc, is a factor of between two and three smaller than its CO(2–1) and CO(3–2) counterparts, which have full sizes of DCO(3 − 2) = 26 ± 0.6 pc and DCO(2 − 1) = 28 ± 0.6 pc, respectively. This result brings into light the many faces of the molecular torus. The torus is connected to the CND through a network of molecular gas streamers detected inside the CND ring. The kinematics of molecular gas show strong departures from circular motions in the torus, the gas streamers, and the CND ring. These velocity field distortions are interconnected and are part of a 3D outflow that reflects the effects of AGN feedback on the kinematics of molecular gas across a wide range of spatial scales around the central engine. In particular, we estimate through modeling that a significant fraction of the gas inside the torus ( ≃ 0.4 − 0.6 × Mtorusgas ≃ 0.4 − 0.6 × M torus gas $ {\simeq}0.4{-}0.6 \times M_{\mathrm{torus}}^{\mathrm{gas}} $ ) and a comparable amount of mass along the gas streamers are outflowing. However, the bulk of the mass, momentum, and energy of the molecular outflow of NGC 1068 is contained at larger radii in the CND region, where the AGN wind and the radio jet are currently pushing the gas assembled at the Inner Lindblad Resonance (ILR) ring of the nuclear stellar bar. Conclusions. In our favored scenario a wide-angle AGN wind launched from the accretion disk of NGC1068 is currently impacting a sizable fraction of the gas inside the torus. However, a large gas reservoir (≃1.2 − 1.8 × 105 M⊙), which lies close to the equatorial plane of the torus, remains unaffected by the feedback of the AGN wind and can therefore continue fueling the AGN for at least ≃1 − 4 Myr. Nevertheless, AGN fueling currently seems thwarted on intermediate scales (15 pc ≤r ≤ 50 pc).
Understanding how super-massive black holes form and grow in the early Universe has become a major challenge
since it was discovered that luminous quasars existed only 700 million years after the Big ...Bang
. Simulations indicate an evolutionary sequence of dust-reddened quasars emerging from heavily dust-obscured starbursts that then transition to unobscured luminous quasars by expelling gas and dust
. Although the last phase has been identified out to a redshift of 7.6 (ref.
), a transitioning quasar has not been found at similar redshifts owing to their faintness at optical and near-infrared wavelengths. Here we report observations of an ultraviolet compact object, GNz7q, associated with a dust-enshrouded starburst at a redshift of 7.1899 ± 0.0005. The host galaxy is more luminous in dust emission than any other known object at this epoch, forming 1,600 solar masses of stars per year within a central radius of 480 parsec. A red point source in the far-ultraviolet is identified in deep, high-resolution imaging and slitless spectroscopy. GNz7q is extremely faint in X-rays, which indicates the emergence of a uniquely ultraviolet compact star-forming region or a Compton-thick super-Eddington black-hole accretion disk at the dusty starburst core. In the latter case, the observed properties are consistent with predictions from cosmological simulations
and suggest that GNz7q is an antecedent to unobscured luminous quasars at later epochs.
Nuclear outflows driven by accreting massive black holes are one of the main feedback mechanisms invoked at high-z to reproduce the distinct separation between star-forming disk galaxies and ...quiescent spheroidal systems. Yet our knowledge of feedback at high-z remains limited by the lack of observations of the multiple gas phases in galaxy outflows. In this work, we use new deep, high spatial resolution ALMA CO(3-2) and archival Very Large Telescope/SINFONI H observations to study the molecular and ionized components of the active galactic nucleus (AGN)-driven outflow in zC400528, a massive main-sequence galaxy at z = 2.3 in the process of quenching. We detect a powerful molecular outflow that shows a positive velocity gradient before a turnover and extends for at least ∼10 kpc from the nuclear region, about three times the projected size of the ionized wind. The molecular gas in the outflow does not reach velocities high enough to escape the galaxy and is therefore expected to be reaccreted. Keeping in mind the various assumptions involved in the analysis, we find that the mass and energetics of the outflow are dominated by the molecular phase. The AGN-driven outflow in zC400528 is powerful enough to deplete the molecular gas reservoir on a timescale comparable to that needed to exhaust it by star formation. This suggests that the nuclear outflow is one of the main quenching engines at work in the observed suppression of the central star formation activity in zC400528.
Context.
Arp220 is the nearest and prototypical ultra-luminous infrared galaxy; it shows evidence of pc-scale molecular outflows in its nuclear regions and strongly perturbed ionised gas kinematics ...on kpc scales. It is therefore an ideal system for investigating outflow mechanisms and feedback phenomena in detail.
Aims.
We investigate the feedback effects on the Arp220 interstellar medium (ISM), deriving a detailed picture of the atomic gas in terms of physical and kinematic properties, with a spatial resolution that had never before been obtained (0.56″, i.e. ∼210 pc).
Methods.
We use optical integral-field spectroscopic observations from VLT/MUSE-AO to obtain spatially resolved stellar and gas kinematics, for both ionised (N II
λ
6583) and neutral (Na ID
λ
λ
5891, 96) components; we also derive dust attenuation, electron density, ionisation conditions, and hydrogen column density maps to characterise the ISM properties.
Results.
Arp220 kinematics reveal the presence of a disturbed kpc-scale disc in the innermost nuclear regions as well as highly perturbed multi-phase (neutral and ionised) gas along the minor axis of the disc, which we interpret as a galactic-scale outflow emerging from the Arp220 eastern nucleus. This outflow involves velocities up to ∼1000 km s
−1
at galactocentric distances of ≈5 kpc; it has a mass rate of ∼50
M
⊙
yr
−1
and kinetic and momentum power of ∼10
43
erg s
−1
and ∼10
35
dyne, respectively. The inferred energetics do not allow us to distinguish the origin of the outflows, namely whether they are active galactic nucleus- or starburst-driven. We also present evidence for enhanced star formation at the edges of – and within – the outflow, with a star-formation rate SFR ∼ 5
M
⊙
yr
−1
(i.e. ∼2% of the total SFR).
Conclusions.
Our findings suggest the presence of powerful winds in Arp220: They might be capable of heating or removing large amounts of gas from the host (“negative feedback”) but could also be responsible for triggering star formation (“positive feedback”).
ABSTRACT
We report the discovery of J0121+0025, an extremely luminous and young star-forming galaxy (MUV = −24.11, log$L_{\rm Ly \alpha } / \rm erg~s^{-1} = 43.8$) at z = 3.244 showing copious Lyman ...continuum (LyC) leakage ($f_{\rm esc, abs} \approx 40{{\ \rm per\ cent}}$). High signal-to-noise ratio rest-frame UV spectroscopy with the Gran Telescopio Canarias reveals a high significance (7.9σ) emission below the Lyman limit (<912 Å), with a flux density level f900 = 0.78 ± 0.10μJy, and strong P-Cygni in wind lines of O vi 1033 Å, N v 1240 Å, and C iv 1550 Å that are indicative of a young age of the starburst (<10 Myr). The spectrum is rich in stellar photospheric features, for which a significant contribution of an AGN at these wavelengths is ruled out. Low-ionization interstellar medium (ISM) absorption lines are also detected, but are weak ($EW_{0} \rm \simeq 1$ Å) and show large residual intensities, suggesting a clumpy geometry of the gas with a non-unity covering fraction or a highly ionized ISM. The contribution of a foreground and AGN contamination to the LyC signal is unlikely. Deep optical to Spitzer/IRAC 4.5 μm imaging show that the spectral energy distribution of J0121+0025 is dominated by the emission of the young starburst, with log($M_{\star }^{\rm burst}/M_{\odot }) = 9.9\pm 0.1$ and $\rm SFR = 981\pm 232$ M⊙ yr−1. J0121+0025 is the most powerful LyC emitter known among the star-forming galaxy population. The discovery of such luminous and young starburst leaking LyC radiation suggests that a significant fraction of LyC photons can escape in sources with a wide range of UV luminosities and are not restricted to the faintest ones as previously thought. These findings might shed further light on the role of luminous starbursts to the cosmic reionization.
We present new CO(2–1) observations of three low-z (d ~350 Mpc) ultra-luminous infrared galaxy (ULIRG) systems (six nuclei) observed with the Atacama large millimeter/submillimeter array (ALMA) at ...high spatial resolution (~500 pc). We detect massive cold molecular gas outflows in five out of six nuclei (Mout ~ (0.3−5) × 108 M⊙). These outflows are spatially resolved with deprojected effective radii between 250 pc and 1 kpc although high-velocity molecular gas is detected up to Rmax ~ 0.5−1.8 kpc (1–6 kpc deprojected). The mass outflow rates are 12–400 M⊙ yr−1 and the inclination corrected average velocity of the outflowing gas is 350–550 km s−1 (vmax = 500−900 km s−1). The origin of these outflows can be explained by the strong nuclear starbursts although the contribution of an obscured active galactic nucleus cannot be completely ruled out. The position angle (PA) of the outflowing gas along the kinematic minor axis of the nuclear molecular disk suggests that the outflow axis is perpendicular to the disk for three of these outflows. Only in one case is the outflow PA clearly not along the kinematic minor axis, which might indicate a different outflow geometry. The outflow depletion times are 15–80 Myr. These are comparable to, although slightly shorter than, the star-formation (SF) depletion times (30–80 Myr). However, we estimate that only 15–30% of the outflowing molecular gas will escape the gravitational potential of the nucleus. The majority of the outflowing gas will return to the disk after 5–10 Myr and become available to form new stars. Therefore, these outflows will not likely completely quench the nuclear starbursts. These star-forming powered molecular outflows would be consistent with being driven by radiation pressure from young stars (i.e., momentum-driven) only if the coupling between radiation and dust increases with increasing SF rates. This can be achieved if the dust optical depth is higher in objects with higher SF. This is the case in at least one of the studied objects. Alternatively, if the outflows are mainly driven by supernovae (SNe), the coupling efficiency between the interstellar medium and SNe must increase with increasing SF levels. The relatively small sizes (<1 kpc) and dynamical times (<3 Myr) of the cold molecular outflows suggests that molecular gas cannot survive longer in the outflow environment or that it cannot form efficiently beyond these distances or times. In addition, the ionized and hot molecular phases have been detected for several of these outflows, so this suggests that outflowing gas can experience phase changes and indicates that the outflowing gas is intrinsically multiphase, likely sharing similar kinematics, but different mass and, therefore, different energy and momentum contributions.