Absorption of high-energy radiation in planetary thermospheres is generally believed to lead to the formation of planetary winds. The resulting mass-loss rates can affect the evolution, particularly ...of small gas planets. We present 1D, spherically symmetric hydrodynamic simulations of the escaping atmospheres of 18 hot gas planets in the solar neighborhood. Our sample only includes strongly irradiated planets, whose expanded atmospheres may be detectable via transit spectroscopy using current instrumentation. The simulations were performed with the PLUTO-CLOUDY interface, which couples a detailed photoionization and plasma simulation code with a general MHD code. We study the thermospheric escape and derive improved estimates for the planetary mass-loss rates. Our simulations reproduce the temperature-pressure profile measured via sodium D absorption in HD 189733 b, but show still unexplained differences in the case of HD 209458 b. In contrast to general assumptions, we find that the gravitationally more tightly bound thermospheres of massive and compact planets, such as HAT-P-2 b are hydrodynamically stable. Compact planets dispose of the radiative energy input through hydrogen Lyα and free-free emission. Radiative cooling is also important in HD 189733 b, but it decreases toward smaller planets like GJ 436 b. Computing the planetary Lyα absorption and emission signals from the simulations, we find that the strong and cool winds of smaller planets mainly cause strong Lyα absorption but little emission. Compact and massive planets with hot, stable thermospheres cause small absorption signals but are strong Lyα emitters, possibly detectable with the current instrumentation. The absorption and emission signals provide a possible distinction between these two classes of thermospheres in hot gas planets. According to our results, WASP-80 and GJ 3470 are currently the most promising targets for observational follow-up aimed at detecting atmospheric Lyα absorption signals.
Energy-limited escape revised Salz, M; Schneider, P C; Czesla, S ...
Astronomy and astrophysics (Berlin),
01/2016, Letnik:
585
Journal Article
Recenzirano
Odprti dostop
Gas planets in close proximity to their host stars experience photoevaporative mass loss. The energy-limited escape concept is generally used to derive estimates for the planetary mass-loss rates. ...Our photoionization hydrodynamics simulations of the thermospheres of hot gas planets show that the energy-limited escape concept is valid only for planets with a gravitational potential lower than log sub(10)(-Phi sub(G)) < 13.11 ergg super(-1) because in these planets the radiative energy input is efficiently used to drive the planetary wind. Massive and compact planets with log sub(10)(-Phi sub(G)) > or =13.6 ergg super(-1) exhibit more tightly bound atmospheres in which the complete radiative energy input is re-emitted through hydrogen Ly alpha and free-free emission. These planets therefore host hydrodynamically stable thermospheres. Between these two extremes the strength of the planetary winds rapidly declines as a result of a decreasing heating efficiency. Small planets undergo enhanced evaporation because they host expanded atmospheres that expose a larger surface to the stellar irradiation. We present scaling laws for the heating efficiency and the expansion radius that depend on the gravitational potential and irradiation level of the planet. The resulting revised energy-limited escape concept can be used to derive estimates for the mass-loss rates of super-Earth-sized planets as well as massive hot Jupiters with hydrogen-dominated atmospheres.
Time Cells in Hippocampal Area CA3 Salz, Daniel M; Tiganj, Zoran; Khasnabish, Srijesa ...
The Journal of neuroscience,
07/2016, Letnik:
36, Številka:
28
Journal Article
Recenzirano
Odprti dostop
Studies on time cells in the hippocampus have so far focused on area CA1 in animals performing memory tasks. Some studies have suggested that temporal processing within the hippocampus may be ...exclusive to CA1 and CA2, but not CA3, and may occur only under strong demands for memory. Here we examined the temporal and spatial coding properties of CA3 and CA1 neurons in rats performing a maze task that demanded working memory and a control task with no explicit working memory demand. In the memory demanding task, CA3 cells exhibited robust temporal modulation similar to the pattern of time cell activity in CA1, and the same populations of cells also exhibited typical place coding patterns in the same task. Furthermore, the temporal and spatial coding patterns of CA1 and CA3 were equivalently robust when animals performed a simplified version of the task that made no demands on working memory. However, time and place coding did differ in that the resolution of temporal coding decreased over time within the delay interval, whereas the resolution of place coding was not systematically affected by distance along the track. These findings support the view that CA1 and CA3 both participate in encoding the temporal and spatial organization of ongoing experience.
Hippocampal "time cells" that fire at specific moments in a temporally structured memory task have so far been observed only in area CA1, and some studies have suggested that temporal coding within the hippocampus is exclusive to CA1. Here we describe time cells also in CA3, and time cells in both areas are observed even without working memory demands, similar to place cells in these areas. However, unlike equivalent spatial coding along a path, temporal coding is nonlinear, with greater temporal resolution earlier than later in temporally structured experiences. These observations reveal both similarities and differences in temporal and spatial coding within the hippocampus of importance to understanding how these features of memory are represented in the hippocampus.
Nonspatial Sequence Coding in CA1 Neurons Allen, Timothy A; Salz, Daniel M; McKenzie, Sam ...
The Journal of neuroscience,
2016-Feb-03, 2016-02-03, 20160203, Letnik:
36, Številka:
5
Journal Article
Recenzirano
Odprti dostop
The hippocampus is critical to the memory for sequences of events, a defining feature of episodic memory. However, the fundamental neuronal mechanisms underlying this capacity remain elusive. While ...considerable research indicates hippocampal neurons can represent sequences of locations, direct evidence of coding for the memory of sequential relationships among nonspatial events remains lacking. To address this important issue, we recorded neural activity in CA1 as rats performed a hippocampus-dependent sequence-memory task. Briefly, the task involves the presentation of repeated sequences of odors at a single port and requires rats to identify each item as "in sequence" or "out of sequence". We report that, while the animals' location and behavior remained constant, hippocampal activity differed depending on the temporal context of items-in this case, whether they were presented in or out of sequence. Some neurons showed this effect across items or sequence positions (general sequence cells), while others exhibited selectivity for specific conjunctions of item and sequence position information (conjunctive sequence cells) or for specific probe types (probe-specific sequence cells). We also found that the temporal context of individual trials could be accurately decoded from the activity of neuronal ensembles, that sequence coding at the single-cell and ensemble level was linked to sequence memory performance, and that slow-gamma oscillations (20-40 Hz) were more strongly modulated by temporal context and performance than theta oscillations (4-12 Hz). These findings provide compelling evidence that sequence coding extends beyond the domain of spatial trajectories and is thus a fundamental function of the hippocampus.
The ability to remember the order of life events depends on the hippocampus, but the underlying neural mechanisms remain poorly understood. Here we addressed this issue by recording neural activity in hippocampal region CA1 while rats performed a nonspatial sequence memory task. We found that hippocampal neurons code for the temporal context of items (whether odors were presented in the correct or incorrect sequential position) and that this activity is linked with memory performance. The discovery of this novel form of temporal coding in hippocampal neurons advances our fundamental understanding of the neurobiology of episodic memory and will serve as a foundation for our cross-species, multitechnique approach aimed at elucidating the neural mechanisms underlying memory impairments in aging and dementia.
Aims. We explore the capabilities of CARMENES for characterising hot-Jupiter atmospheres by targeting multiple water bands, in particular, those at 1.15 and 1.4 μm. Hubble Space Telescope ...observations suggest that this wavelength region is relevant for distinguishing between hazy and/or cloudy and clear atmospheres. Methods. We observed one transit of the hot Jupiter HD 189733 b with CARMENES. Telluric and stellar absorption lines were removed using SYSREM, which performs a principal component analysis including proper error propagation. The residual spectra were analysed for water absorption with cross-correlation techniques using synthetic atmospheric absorption models. Results. We report a cross-correlation peak at a signal-to-noise ratio (S/N) of 6.6, revealing the presence of water in the transmission spectrum of HD 189733 b. The absorption signal appeared slightly blueshifted at –3.9 ± 1.3 km s−1. We measured the individual cross-correlation signals of the water bands at 1.15 and 1.4 μm, finding cross-correlation peaks at S/N of 4.9 and 4.4, respectively. The 1.4 μm feature is consistent with that observed with the Hubble Space Telescope. Conclusions. The water bands studied in this work have been mainly observed in a handful of planets from space. Being able also to detect them individually from the ground at higher spectral resolution can provide insightful information to constrain the properties of exoplanet atmospheres. Although the current multi-band detections can not yet constrain atmospheric haze models for HD 189733 b, future observations at higher S/N could provide an alternative way to achieve this aim.
We present three transit observations of HD 189733 b obtained with the high-resolution spectrograph CARMENES at Calar Alto. A strong absorption signal is detected in the near-infrared He I triplet at ...10830 Å in all three transits. During mid-transit, the mean absorption level is 0.88 ± 0.04% measured in a ±10 km s−1 range at a net blueshift of − 3.5 ± 0.4 km s−1 (10829.84–10830.57 Å). The absorption signal exhibits radial velocities of + 6.5 ± 3.1 km s−1 and − 12.6 ± 1.0 km s−1 during ingress and egress, respectively; all radial velocities are measured in the planetary rest frame. We show that stellar activity related pseudo-signals interfere with the planetary atmospheric absorption signal. They could contribute as much as 80% of the observed signal and might also affect the observed radial velocity signature, but pseudo-signals are very unlikely to explain the entire signal. The observed line ratio between the two unresolved and the third line of the He I triplet is 2.8 ± 0.2, which strongly deviates from the value expected for an optically thin atmospheres. When interpreted in terms of absorption in the planetary atmosphere, this favors a compact helium atmosphere with an extent of only 0.2 planetary radii and a substantial column density on the order of 4 × 1012 cm−2. The observed radial velocities can be understood either in terms of atmospheric circulation with equatorial superrotation or as a sign of an asymmetric atmospheric component of evaporating material. We detect no clear signature of ongoing evaporation, like pre- or post-transit absorption, which could indicate material beyond the planetary Roche lobe, or radial velocities in excess of the escape velocity. These findings do not contradict planetary evaporation, but only show that the detected helium absorption in HD 189733 b does not trace the atmospheric layers that show pronounced escape signatures.
Close-in gas planets are subject to continuous photoevaporation that can erode their volatile envelopes. Today, ongoing mass loss has been confirmed in a few individual systems via transit ...observations in the ultraviolet spectral range. We demonstrate that the Ultraviolet/Optical Telescope (UVOT) onboard the Neil Gehrels Swift Observatory enables photometry to a relative accuracy of about 0.5% and present the first near-UV (200–270 nm, NUV) transit observations of WASP-121 b, a hot Jupiter with one of the highest predicted mass-loss rates. The data cover the orbital phases 0.85–1.15 with three visits. We measure a broadband NUV transit depth of 2.10 ± 0.29%. While still consistent with the optical value of 1.55%, the NUV data indicate excess absorption of 0.55% at a 1.9σ level. Such excess absorption is known from the WASP-12 system, and both of these hot Jupiters are expected to undergo mass loss at extremely high rates. With a Cloudy simulation, we show that absorption lines of Fe II in a dense extended atmosphere can cause broadband near-UV absorption at the 0.5% level. Given the numerous lines of low-ionization metals, the NUV range is a promising tracer of photoevaporation in the hottest gas planets.
Context.
HD 209458 b is an exoplanet with an upper atmosphere undergoing blow-off escape that has mainly been studied using measurements of the Ly
α
absorption. Recently, high-resolution measurements ...of absorption in the He
I
triplet line at 10 830 Å of several exoplanets (including HD 209458 b) have been reported, creating a new opportunity to probe escaping atmospheres.
Aims.
We aim to better understand the atmospheric regions of HD 209458 b from where the escape originates.
Methods.
We developed a 1D hydrodynamic model with spherical symmetry for the HD 209458 b thermosphere coupled with a non-local thermodynamic model for the population of the He
I
triplet state. In addition, we performed high-resolution radiative transfer calculations of synthetic spectra for the helium triplet lines and compared them with the measured absorption spectrum in order to retrieve information about the atmospheric parameters.
Results.
We find that the measured spectrum constrains the H/H
+
transition altitude occurring in the range of 1.2
R
P
–1.9
R
P
. Hydrogen is almost fully ionised at altitudes above 2.9
R
P
. We also find that the X-ray and extreme ultraviolet absorption takes place at effective radii from 1.16 to 1.30
R
P
, and that the He
I
triplet peak density occurs at altitudes from 1.04 to 1.60
R
P
. Additionally, the averaged mean molecular weight is confined to the 0.61–0.73 g mole
−1
interval, and the thermospheric H/He ratio should be larger than 90/10, and most likely approximately 98/2. We also provide a one-to-one relationship between mass-loss rate and temperature. Based on the energy-limited escape approach and assuming heating efficiencies of 0.1–0.2, we find a mass-loss rate in the range of (0.42–1.00) ×10
11
g s
−1
and a corresponding temperature range of 7125–8125 K.
Conclusions.
The analysis of the measured He
I
triplet absorption spectrum significantly constrains the thermospheric structure of HD 209458 b and advances our knowledge of its escaping atmosphere.
Alcohol Use Disorder (AUD) has been associated with abnormalities in hippocampal volumes, but these relationships have not been fully explored with respect to sub-regional volumes, nor in association ...with individual characteristics such as age, gender differences, drinking history, and memory. The present study examined the impact of those variables in relation to hippocampal subfield volumes in abstinent men and women with a history of AUD. Using Magnetic Resonance Imaging at 3 Tesla, we obtained brain images from 67 participants with AUD (31 women) and 64 nonalcoholic control (NC) participants (31 women). The average duration of the most recent period of sobriety for AUD participants was 7.1 years. We used Freesurfer 6.0 to segment the hippocampus into 12 regions. These were imputed into statistical models to examine the relationships of brain volume with AUD group, age, gender, memory, and drinking history. Interactions with gender and age were of particular interest. Compared to the NC group, the AUD group had approximately 5% smaller subiculum, CA1, molecular layer, and hippocampal tail regions. Age was negatively associated with volumes for the AUD group in the subiculum and the hippocampal tail, but no significant interactions with gender were identified. The relationships for delayed and immediate memory with hippocampal tail volume differed for AUD and NC groups: Higher scores on tests of immediate and delayed memory were associated with smaller volumes in the AUD group, but larger volumes in the NC group. Length of sobriety was associated with decreasing CA1 volume in women (0.19% per year) and increasing volume size in men (0.38% per year). The course of abstinence on CA1 volume differed for men and women, and the differential relationships of subfield volumes to age and memory could indicate a distinction in the impact of AUD on functions of the hippocampal tail. These findings confirm and extend evidence that AUD, age, gender, memory, and abstinence differentially impact volumes of component parts of the hippocampus.
Aims. We aim at detecting water vapor in the atmosphere of the hot Jupiter HD 209458 b and perform a multi-band study in the near infrared with CARMENES. Methods. The water vapor absorption lines ...from the atmosphere of the planet are Doppler-shifted due to the large change in its radial velocity during transit. This shift is of the order of tens of km s−1, whilst the Earth’s telluric and the stellar lines can be considered quasi-static. We took advantage of this shift to remove the telluric and stellar lines using SYSREM, which performs a principal component analysis including proper error propagation. The residual spectra contain the signal from thousands of planetary molecular lines well below the noise level. We retrieve the information from those lines by cross-correlating the residual spectra with models of the atmospheric absorption of the planet. Results. We find a cross-correlation signal with a signal-to-noise ratio (S/N) of 6.4, revealing H2O in HD 209458 b. We obtain a net blueshift of the signal of –5.2 −1.3+2.6 $^{+2.6}_{-1.3}$ −1.3+2.6 km s−1 that, despite the large error bars, is a firm indication of day- to night-side winds at the terminator of this hot Jupiter. Additionally, we performed a multi-band study for the detection of H2O individually from the three near infrared bands covered by CARMENES. We detect H2O from its 0.96–1.06 μm band with a S/N of 5.8, and also find hints of a detection from the 1.06–1.26 μm band, with a low S/N of 2.8. No clear planetary signal is found from the 1.26–1.62 μm band. Conclusions. Our significant H2O signal at 0.96–1.06 μm in HD 209458 b represents the first detection of H2O from this band individually, the bluest one to date. The unfavorable observational conditions might be the reason for the inconclusive detection from the stronger 1.15 and 1.4 μm bands. H2O is detected from the 0.96–1.06 μm band in HD 209458 b, but hardly in HD 189733 b, which supports a stronger aerosol extinction in the latter, in line with previous studies. Future data gathered at more stable conditions and with larger S/N at both optical and near-infrared wavelengths could help to characterize the presence of aerosols in HD 209458 b and other planets.
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