Mobile ad hoc networks (MANETs) have gained a lot of interests in research communities for the infrastructure-less self-organizing nature. A MANET with fleet cyclists using smartphones forms a two-tier mobile long-thin network (MLTN) along a common cycling route, where the high-tier network is composed of 3G/LTE interfaces and the low-tier network is composed of IEEE 802.11 interfaces. The low-tier network may consist of several path-like networks. This work investigates cooperative sensing data collection and distribution with packet collision avoidance in a two-tier MLTN. As numbers of cyclists upload their sensing data and download global fleet information frequently, serious bandwidth and latency problems may result if all members rely on their high-tier interfaces. We designed and analyzed a cooperative framework consisting of a distributed grouping mechanism, a group merging and splitting method, and a sensing data aggregation scheme. Through cooperation between the two tiers, the proposed framework outperforms existing works by significantly reducing the 3G/LTE data transmission and the number of 3G/LTE connections.