Mechanical site preparation (MSP) is used prior to planting to control competing vegetation and enhance soil conditions, particularly in areas prone to paludification. Tree planting density can be ...adapted to the management context and objectives, as it influences yield and wood quality. However, the combined effects of MSP and planting density on understory vegetation composition, functional traits, and diversity remain uncertain. We thus conducted a study in the Clay Belt region of northwestern Quebec, Canada. After careful logging, the study area was divided into nine sites, each receiving one of three treatments: plowing, disc trenching, or no preparation. Sites were further divided into two, with black spruce (Picea mariana Mill. Britton, Sterns & Poggenb.) seedlings planted at either a low planting density of 1100 seedlings ha-1 or a high planting density of 2500 seedlings ha-1. After nine years, we assessed understory composition, diversity, key functional traits, sapling density and growth of planted trees. Careful logging alone led to a higher density of naturally established conifers compared to plowing or disc trenching. The interaction between planting density and MSP significantly influenced understory diversity and composition in plowed plots. Understory composition was affected by the soil C/N ratio, coniferous species, and deciduous species density. The growth of black spruce was notably enhanced with higher planting density in the plow treatment only. Neither planting density nor MSP alone affected tree height and diameter. Our results suggest that combining plowing with high-density planting can enhance stand growth and improve forest productivity. These findings guide future research on paludified forests.
In boreal forests in Canada, broadleaf stands are characterized by generally well-drained soils and a humus-rich layer. In contrast, spruce-moss stands are often characterized by more poorly drained ...soils and acidic humus layer. However, presence of these two forest types in various degrees of mixture in stands can be beneficial to spruce seedlings productivity. It was hypothesized that leaf litter and humus from pure spruces-moss stands, pure broadleaf stands, and mixed stand may influence Black spruce (Picea mariana (Mill.) BSP) seedling growth and development differently. A greenhouse experiment was carried out to evaluate the effect of different leaf litter and different humus on spruces seedlings. Our results suggest better development for seedlings grown in humus from mixed stands and pure broadleaf stands compared to humus from pure B. spruce or standard forest nursery substrate. Furthermore, leaf litter from broadleaf trees species, such as species Speckle alder (Alnus rugosa (Du Roi) R.T. Clausen), T. aspen (Populus tremuloides Michx), Willows (Salix spp.) and Paper birch (Betula papyrifera Marsh.), has shown distinct results in the growth and development of B. spruce seedlings in greenhouse. Furthermore, promotion of mixed stand can increase B. spruce productivity by improving the physicochemical composition of the forest floor.
Forest vegetation management can improve planted seedling survival and growth and is thus widely used in plantation silviculture. In some jurisdictions, mechanical release using brushsaws has ...replaced the traditional use of chemical herbicides for forest vegetation management purposes. However, its associated costs and the increasing difficulty of finding qualified labor represent a challenge. The browsing of competition by large herbivores may represent an alternative to mechanical release when planted seedlings are resistant to browsing. Here, we compare the efficacy of moose browsing relative to mechanical release in controlling competing vegetation and in promoting white spruce growth in plantations. In a high moose density region, we used an experimental design consisting of four pairs of moose exclosures and unfenced plots; fifty percent of both the access-restricted and unrestricted study areas received a mechanical release treatment. Moose browsing was more efficient than mechanical release in diminishing the sapling density and basal area of competing species. Mechanical release only reduced the sapling density of taller competitors (height > 201 cm), whereas browsing reduced the sapling densities of competitors across a greater size range (height > 130 cm). These effects of moose browsing on competition translated into a greater positive effect of moose browsing on the basal area of planted spruces. We attribute the higher effectiveness of moose browsing relative to mechanical release to its chronic nature. Moose browsed continuously throughout the year and for multiple years, whereas mechanical release was applied only one time between the second and fourth years after planting. Our results suggest that pairing wildlife management and silviculture decisions could be in the best interest of both the hunting and forestry industries in regions where plantations are frequent and use browse-resistant crop trees. Favouring browsers in controlling the density of competing species could increase the hunting experience and income, while providing an effective, cost-free, and socially acceptable forest vegetation management service.
In the province of Québec (Canada), pre-commercial thinning (PCT) is a common silvicultural practice applied to young black spruce (Picea mariana (Mill.) BSP) stands. PCT removes some of the ...competing vegetation and smaller black spruce stems, in order to improve growth rates and reduce forest rotation intervals. It is uncertain whether this positive response in black spruce growth is primarily due to lower competition for resources or to other mechanisms, which may vary according to climate or edaphic conditions. We sampled soils and black spruce needles in PCT-treated and non-treated control plots occurring in two climate regimes, as well as on two contrasting soil parent materials within one of these two climate regimes (i.e., three “site types”). We performed our sampling approximately 20 years after treatment. Paired treatment plots (i.e., PCT vs. control) were replicated at four independent sites in each of the three site types, for a total of 24 plots. Over two consecutive years, we measured stand structural characteristics, indices of soil N fertility, soil microbial activity, indices of soil moisture availability, canopy openness, and foliar characteristics in each plot. In each site type, PCT decreased total basal area but increased radial growth of individual trees. Across all plots, soil N mineralization rates measured in 2016 were positively related to foliar N concentrations of one-year-old needles collected in 2017. Annual precipitation, drainage class, potential evapotranspiration, and climate moisture index all indicated that plots occurring in the drier climate and on glacial till deposits were more prone to summer moisture deficits. Accordingly, PCT increased forest floor moisture only in this site type, which may benefit tree growth. In the wetter climate and on poorly drained soils, however, we found evidence that PCT reduces soil N fertility, presumably by increasing the spread of ericaceous shrubs in the understory. In the dry fertile site type, the range in canopy openness was substantially higher (12–37%) and correlated negatively with tree diameter, suggesting that greater light availability did not improve tree growth. Taken collectively, our data suggest that PCT increases black spruce growth across a broad range of site conditions found in Québec, presumably by lowering intraspecific competition for resources. However, on drier sites, PCT may also benefit trees by increasing soil moisture availability, whereas wetter climates may mitigate the beneficial effect of PCT due to a loss of soil N fertility.
We evaluated the effectiveness of commercial thinning mainly from below (CT; 0, 26%, 32% and 40% merchantable basal area removals) in meeting wood production demands and developing structural ...heterogeneity in a balsam fir (Abies balsamea (L.) Mill) and spruce (Picea spp.) stand. After 10 years, 32%-40% removals showed a 12%-18% increase in mean diameter and 27%-38% increase in gross merchantable volume (GMV) per tree compared to the unthinned control. At the stand level, all thinning treatments generated as much cumulative GMV (harvested volume + GMV after 10 years) and gross sawlog volume per hectare as the unthinned control. As for stand structure, eight out of nine thinned experimental units showed increased structural heterogeneity after 10 years, i.e., irregular, positively-skewed diameter distribution with an elongated right tail toward larger trees. The diameter distribution in the unthinned control became more symmetric, unimodal and regular over time, with fewer saplings than at the beginning of the experiment and lower density of larger trees compared to CT. Regeneration density and stocking were abundant in all treatments, largely dominated by balsam fir. Results indicate that thinning can be used to meet wood production objectives and help develop structural heterogeneity in this forest.
Forest plantations play an increasingly important role in meeting global demand for wood. They usually have higher yield than naturally regenerated forests. Thus, plantations can support economically ...viable wood production, enable forest conservation elsewere, help mitigate climate change by contributing to carbon sequestration and increase forest resilience and resistance to biotic and abiotic stressors. If yield of plantations is not as high as anticipated, then their use could generate important sustainability issues. There are still major gaps in our understanding of the factors that influence yield, even with respect to black spruce, white spruce, and jack pine, three of the most commonly planted tree species in northeastern North America. Our objective was to evaluate the yield of forest plantations of these species over a 416 000 km
2
region that was representative of northeastern North American forests. Contrary to our prediction, realized yield of operational plantations was consistently lower than anticipated. Site index and competition both played a significant role in determining the yield of plantations. In the context of uncertain realized yield of operational plantations, we emphasize the necessity of relying on adaptive management to determine harvest levels that are compatible with sustainable management objectives.
•Resilience of natural forests can jeopardize or enhance plantation productivity.•Unproductive plantations will likely generate sustainability issues.•Plantation scenarios should aim toward ...resilience-driven successional trajectories.
The demand for wood products continues to increase globally. Productivity of forest plantations can be greater than that of naturally regenerated forests. Plantation forestry could thus be employed to meet up to 75% of global wood supply by 2050. The resilience of natural forests could jeopardize plantation productivity. If the plantation scenario is not oriented in the same direction as the natural successional trajectories that are driven by resilience, then naturally regenerating tree species could recover to the detriment of planted species. Unproductive plantations will likely generate sustainability issues, as they must provide ecosystem services (e.g., wood fiber production) while being economically viable for forest management to be sustainable. Our general objective of our study was to assess whether resilience of natural forests can jeopardized plantation productivity. We studied successional trajectories in recent, young and old black spruce plantations that are located within the balsam fir ecological region on balsam fir ecological site types. To compare with a plantation scenario that was more susceptible to aiming in the same direction as resilience-driven successional trajectories, we also studied successional trajectories in recent, young and old black spruce plantations that are not only located within the balsam fir ecological region, but on black spruce ecological site types. Successional trajectories on balsam fir ecological site types pointed towards recovery of balsam fir and white birch, to the detriment of black spruce. Successional trajectories on black spruce ecological site types pointed towards the recovery of black spruce. Thus, we showed that resilience of natural forests could jeopardize plantation productivity when the plantation scenario is not oriented in the same direction as resilience-driven successional trajectories. It also can enhance plantation productivity when the regime is oriented in the same direction as resilience-driven successional trajectories. To ensure that plantations are economically viable and promote sustainability, forest managers should thus favor plantation scenarios that point in the same direction as resilience-driven successional trajectories. Finally, we suggest adopting such scenarios, and stress that it would also be necessary to develop regimes that would promote sustainability in the context of alternative successional trajectories that will undoubtedly arise because of global changes.
Clear-cutting is used by foresters worldwide to harvest timber from forest ecosystems. Clear-cutting of cool–wet boreal forests in late successional stages can maintain successional trajectories ...pointed toward the composition of a late successional stages. However, clear-cutting of warm–dry boreal forests in late successional stages can trigger reversion of successional trajectories back toward forests of earlier stages because early-successional shade-intolerant species are more abundant than in cool–wet boreal forests, a process that is referred to as a successional setback. Such successional setbacks can generate sustainability issues by extending the expected time to compositional recovery after clear-cutting. This can lead to an overestimation of allowable cuts of economically important late-successional species and subsequently to a temporary forest composition conversion if the occurrence of successional setbacks remains unassessed. Temperate forests in late successional stages are warmer and drier than boreal forests and consequently include more early-successional shade-intolerant species susceptible to encroach clear-cut areas. Even if current ecological knowledge suggests that temperate forests could be susceptible to post-clearcut successional setbacks, a comprehensive assessment has yet to be undertaken. The objective of the present study was to assess whether clear-cutting has triggered successional setbacks in temperate forests. Therefore, we studied post-clearcut successional trajectories by using forest inventory data covering the entire temperate forest of the province of Québec, eastern Canada (209 000 km2). Clear-cutting triggered successional setbacks in both ecological regions forming the temperate forest. After clear-cutting, successional trajectories of trees pointed toward the composition of an early successional stage. To address this sustainable management issue in a comprehensive manner with clear-cutting, foresters could use partial cut approaches.
•Clear-cutting can trigger successional setbacks in temperate forests.•Such setbacks could extend compositional recovery by an additional century.•This can lead to overestimation of allowable cuts of late-successional species.•With clear-cutting, foresters could use partial cutting to answer this issue.
•Boreal forests experience the effects of climate change faster than any other biome.•Thinning increases residual tree growth, particularly on moisture-limited sites.•On warm-dry sites, greater water ...availability increases radial tree growth.•On wet sites, greater water availability decreases the radial growth.•Black spruce growth may benefit from future warming on relatively wet sites.
Rising temperatures are likely to increase the risk of drought across the globe over the next century. Boreal forests are particularly vulnerable to drought because temperatures within these biomes are projected to warm the fastest. Warm and dry conditions can reduce tree growth, particularly in regions that are already moisture-limited, which may reduce forest productivity. Forest stand density management, such as pre-commercial thinning (PCT), can reduce moisture stress for residual trees by reducing canopy evaporation and increasing soil water availability. PCT is applied 15–20 years after clearcut and removes smaller stems to increase resource availability to residual trees, thereby increasing diameter growth and wood value. How PCT can mitigate climate change impact in boreal forests is still unclear, partly due to heterogeneity in moisture availability and growth response. This study tests the relative effect of PCT on radial growth of black spruce (Picea mariana (Mill.) B.S.P.) on sites with varying water availability to inform forest managers about PCT’s potential to mitigate anticipated effects of drought on tree growth. Tree discs from PCT and non-thinned plots were harvested from three contrasting site types in eastern Canada (warm-dry, warm-wet and cool wet). Using dendrochronology, the relationship between annual ring width index (RWI) and standardized-evapotranspiration index (SPEI) was explored in the time since PCT and during known climate anomalies. RWI showed a positive growth response to increasing SPEI on warm-dry sites, but a negative growth response on warm-wet and cool-wet sites. PCT provided a greater benefit to radial growth on warm-dry site types in the 15 years since treatment but provided no additional benefits during years of climate anomalies. Results suggest that PCT will remain an important forest management practice on moisture-limited sites in order to maintain black spruce productivity, whereas tree growth may benefit from future warming on relatively wet sites.
•Both functional diversity and community weighted trait mean affected tree growth.•Improving understory functional richness promote tree growth in clear-cut forests.•Functional evenness or divergence ...matters for tree growth in careful logged forests.•Morphological, regeneration or resource utilization traits can impact tree growth.•Identifying disturbance threshold that can activate more facilitation is suggested.
Understory vegetation plays an important role in maintaining biodiversity, soil nutrient cycling and carbon stocks, yet the ability of understory functional traits to predict forest productivity after harvesting disturbance is unknown. Our objective was to investigate the utility of individual traits (the community-weighted trait mean) and combined information from multiple traits (functional diversity indices) of understory plants in predicting tree diameter growth of black spruce (Picea mariana (Mill.) B.S.P.) after two harvesting methods (clearcutting and careful logging). We used vegetation and tree growth data from a network of field plots (171 plots on 43 sites) established in black spruce dominated forests across Ontario and Quebec, in northeastern Canada. Functional traits (11 traits) of dominant understory species that reflect plant morphology, regeneration strategy, and resource utilization were collected. We then used generalized linear mixed models (GLMMs) to model the response of tree diameter increment to functional diversity indices, community-weighted trait means and their interactions with harvesting method. Our results showed that: 1) at the functional diversity level, functional richness had a positive relationship with tree growth in clear-cut forests, while functional evenness showed a negative and functional divergence a positive relationship with tree growth in careful logging forests; 2) at the individual trait level, understory species with relatively large morphological trait values often promoted tree growth in both forest types, with stronger relationships in careful logging than in clear-cut forests. For regeneration traits, heavier seed weight or permanent seed bank persistence of understory plants had negative relationships with tree growth, and those relationships were either only found in careful logging forests or did not depend on harvesting type. A positive relationship was found between vegetative propagation species and tree growth, which was stronger in careful logging than in clear-cut forests. Species preferring either humid or xeric habitat had positive relationships with tree growth; both shade-tolerant and shade-intolerant species also positively correlated to tree growth. In conclusion, our study shows the important roles of understory traits in predicting tree growth based on both niche complementarity and mass-ratio hypotheses. Moreover, disturbance severity influenced the balance or degree of the positive and negative interactions that occurred between understory plants and black spruce tree growth. We suggest that identifying the threshold of harvesting disturbance severity that can activate facilitation interactions between understory plants and crop trees would be an important topic for future study.
PDF
