Morphological analyses of three glomoid spore-producing fungi suggested that two of them were undescribed species of Glomeraceae (phylum Glomeromycota ), and the third differed slightly from Dominikia glomerocarpica and Epigeocarpum crypticum , recently described in Glomeraceae . The first two fungi originated from the Mediterranean Sea sand dunes of the Peloponnese, Greece, and the third was originally found in a tree plantation in Yokohama City, Japan. Phylogenetic analyses of sequences of the 45S nuc rDNA region and the RPB1 gene showed that (i) the three fungi belonged to Glomeraceae ; (ii) the first two represented a new genus, here described as Complexispora gen. nov. with C. multistratosa sp. nov. and C. mediterranea sp. nov. and (iii) the third enlarged the monospecific genus Epigeocarpum , as E. japonicum sp. nov.