This paper presents an updated checklist of the butterflies of Europe, together with their original name combinations, and their occurrence status in each European country. According to this ...checklist, 496 species of the superfamily Papilionoidea occur in Europe. Changes in comparison with the last version (2.6.2) of Fauna Europaea are discussed. Compared to that version, 16 species are new additions, either due to cryptic species most of which have been discovered by molecular methods (13 cases) or due to discoveries of Asian species on the eastern border of the European territory in the Ural mountains (three cases). On the other hand, nine species had to be removed from the list, because they either do not occur in Europe or lost their species status due to new evidence. In addition, three species names had to be changed and 30 species changed their combination due to new evidence on phylogenetic relationships. Furthermore, minor corrections were applied to some authors’ names and years of publication. Finally, the name
Polyommatusottomanus
Lefèbvre, 1831, which is threatened by its senior synonym
Lycaenalegeri
Freyer, 1830, is declared a
nomen protectum
, thereby conserving its name in the current combination
Lycaenaottomana
.
Abstract
Here, we present the largest, global dataset of Lepidopteran traits, focusing initially on butterflies (
ca
. 12,500 species records). These traits are derived from field guides, taxonomic ...treatments, and other literature resources. We present traits on wing size, phenology,voltinism, diapause/overwintering stage, hostplant associations, and habitat affinities (canopy, edge, moisture, and disturbance). This dataset will facilitate comparative research on butterfly ecology and evolution and our goal is to inspire future research collaboration and the continued development of this dataset.
Two new species and two new subspecies of the satyrine butterfly genus Punargentus Heimlich, P. atusparia sp. n., P. heimlichi sp. n., P. blanchardi libertas ssp. n. and P. atusparia yupania ssp. n., ...are described from north-central Peru (Ancash, Huánuco and La Libertad). The affinities of the new taxa are evaluated based on molecular data obtained using target enrichment and COI barcoding. Punargentus is fully supported and consists of two branches, the P. lamna clade with four species and P. blanchardi clade with three species, two of which are described here. Species in the P. blanchardi clade occur in Andean shrubland, at altitudes between 2800–3600 m, whereas those in the P. lamna clade occur in puna grassland at 3600–4600 m, with occasional overlaps. Based on their habitat preferences, none of the newly described species are in need of immediate conservation measures. The genus Etcheverrius Herrera is reinstated, with E. chiliensis (Guérin-Méneville) and E. tandilensis (Köhler), comb. n.
Comparing specimens curated in MfN as primary types of Hesperiidae names proposed by Carl Plötz with the original descriptions and unpublished drawings reveals a number of inconsistencies that we ...address. Lectotypes are designated for Telegonus labriaris Butler, 1877, Eudamus jalapus Plötz, 1881, and Apaustus interpunctata Plötz, 1884. Neotypes are designated for Netrocoryne seneca Plötz, 1882 and Hesperia irma Plötz, 1882. Hesperia ulphila Plötz, 1883 is treated as a nomen dubium. As a result of these designations, the following are junior objective synonyms: Netrocoryne seneca Plötz, 1882 of Telegonus labriaris Butler, 1877 and Hesperia irma Plötz, 1882 of Pamphila irma Möschler, 1879, the latter two names being homonyms. In all these instances, the original descriptions are satisfied, and, except for A. interpunctata, the current application of these names is unaffected. Taxonomically, Callimormus diaeses Schaus, 1902, stat. rest. is a valid species, not a synonym of A. interpunctata, which is a junior subjective synonym of Eutocus vetulus (Mabille, 1883).
Talides basistrigata Eaton, 1932, originalmente descrita de Perú, posteriormente fue transferida por Evans (1955) al género Hylephila Billberg, 1820, y considerada como una subespecie de H. phyleus ...(Drury, 1773), argumentando que basistrigata ocurre en ambientes más secos que los que habita phyleus. Actualmente se mantiene el rango taxonómico propuesto por Evans. El objetivo del presente trabajo es restituir el estatus taxonómico de especie válida para basistrigata con base en un análisis integrativo considerando aspectos morfológicos, ecológicos y genéticos. Se examinó las estructuras genitálicas en ejemplares de basistrigata y phyleus, se obtuvo secuencias del gen mitocondrial COI para ambas especies y se recopiló datos en campo para conocer su comportamiento de oviposición en los alrededores de Arequipa, suroeste de Perú. Como resultado del examen morfológico se halló diferencias notorias entre las estructuras genitálicas de ambas especies, principalmente en los procesos caudales de las valvas, la disposición de las pectinas y la forma del uncus en los machos. Con relación al análisis genético, se halló una distancia genética mayor a 3% entre ambas especies, apoyando así el estatus taxonómico de especie para basistrigata. Adicionalmente, desde una perspectiva ecológica, se registró que ambas especies utilizan distintos mecanismos de oviposición y diferentes plantas hospederas, exhibiendo además simpatría espacial y sincronía temporal en la vertiente occidental de los Andes en Perú. Se redescribe el macho y hembra adultos de basistrigata y se designa su lectotipo.
The systematics of the enigmatic and rather uncommon species of Sea Hayward and Cybdelis Boisduval is assessed through morphological and molecular analyses. The aim of this paper is three‐fold: (i) ...to ascertain the phylogenetic position of Sea and Cybdelis in Biblidinae; (ii) to determine the evolutionary relationships among species of Sea and Cybdelis; and (iii) to review the taxonomy of Sea and Cybdelis. The result of the molecular analysis recovers Sea sister to Cybdelis, and this clade sister to Eubagina (sensu Lamas, ), rendering Epicaliina (sensu Lamas, ) paraphyletic. A new tribe, Cybdelini trib.n. is erected to include Sea and Cybdelis; the other genera formerly included in Epicaliina (sensu Lamas, ), Eunica Hübner and Sevenia Koçak are assigned to Eunicini trib.rest., and the remaining genera of Biblidinae to other previously recognized tribes, similar to those implicitly proposed by Wahlberg et al. (). The morphological analysis recovers similar results and the following relationships among species of Cybdelis: C. phaesyla (Hübner) (C. boliviana Salvin (C. mnasylus Doubleday (C. petronita Burmeister stat.rest. + C. thrasylla C. Felder & R. Felder stat.rest.))). Sea and Cybdelis are recognized as distinct genera; a new subspecies, S. sophronia jenkinsi Dias, Siewert & Freitas ssp.n., distributed from Venezuela to Bolivia along the Andes eastern foothills, is described; and C. petronita and C. thrasylla are accepted as valid species. Lectotypes for C. boliviana, C. peruviana Staudinger, C. mnasylus, C. thrasylla, C. petronita and C. mnasylus var. meridionalis Hall and a neotype for Vanessa sophronia Godart are designated. Additionally, dichotomous keys and distribution maps to all taxa are provided; male genitalia and hypandria are illustrated for all species; generic characters of the head, labial palpi and legs are illustrated; immature stages of some species also are figured.
The published work has been registered on ZooBank:
http://zoobank.org/urn:lsid:zoobank.org:pub:D8EBB649‐5E9D‐4218‐B1F1‐7A69DA775FD0.
Molecular analysis recovers the validity of the sister genera Sea and Cybdelis, and this clade as sister to Eubagina, rendering Epicaliina (sensu Lamas, ) paraphyletic.
Cybdelini trib.n. includes Sea and Cybdelis, other genera formerly in Epicaliina (sensu Lamas, ) are assigned to Eunicini trib.rest., and the remaining genera to other previously recognized tribes.
Morphologic analysis recovers the relationship among species of Sea and Cybdelis; two subspecies of Cybdelis are restituted to species and a new subspecies of Sea sophronia is described.
Sepona Freitas and Barbosa, gen. nov. is proposed for the Neotropical satyrine butterfly species Euptychia punctataWeymer, 1911 and its junior subjective synonyms Euptychia griseolaWeymer, 1911 and ...Taygetis indecisa Ribeiro, 1931. The new genus has a distinctive wing pattern and shape of the valvae in the male genitalia, the latter being a unique autapomorphy within the subtribe Euptychiina. Based on molecular data, this genus is not sister to any other single euptychiine genus, instead appearing as the sister to all remaining genera in the Taygetis clade. The present paper illustrates the complexity of the taxonomy of Euptychiina, and the importance of using different sources of evidence in taxonomic studies.
This paper presents an updated checklist of the butterflies of Europe, together with their original name combinations, and their occurrence status in each European country. According to this ...checklist, 496 species of the superfamily Papilionoidea occur in Europe. Changes in comparison with the last version (2.6.2) of Fauna Europaea are discussed. Compared to that version, 16 species are new additions, either due to cryptic species most of which have been discovered by molecular methods (13 cases) or due to discoveries of Asian species on the eastern border of the European territory in the Ural mountains (three cases). On the other hand, nine species had to be removed from the list, because they either do not occur in Europe or lost their species status due to new evidence. In addition, three species names had to be changed and 30 species changed their combination due to new evidence on phylogenetic relationships. Furthermore, minor corrections were applied to some authors' names and years of publication. Finally, the name
Lefèbvre, 1831, which is threatened by its senior synonym
Freyer, 1830, is declared a
, thereby conserving its name in the current combination
.
'Suture zones' are areas where hybrid and contact zones of multiple taxa are clustered. Such zones have been regarded as strong evidence for allopatric divergence by proponents of the Pleistocene ...forest refugia theory, a vicariance hypothesis frequently used to explain diversification in the Amazon basin. A central prediction of the refugia and other vicariance theories is that the taxa should have a common history so that divergence times should be coincident among taxa. A suture zone for Ithomiinae butterflies near Tarapoto, NE Peru, was therefore studied to examine divergence times of taxa in contact across the zone. We sequenced 1619 bp of the mitochondrial COI/COII region in 172 individuals of 31 species from across the suture zone. Inferred divergence times differed remarkably, with divergence between some pairs of widespread species (each of which may have two or more subspecies interacting in the zone, as in the genus Melinaea) being considerably less than that between hybridizing subspecies in other genera (for instance in Oleria). Our data therefore strongly refute a simple hypothesis of simultaneous vicariance and suggest that ongoing parapatric or other modes of differentiation in continuous forest may be important in driving diversification in Amazonia.
Nomenclature, far from being a dry dusty subject, is today more relevant than ever before. Researchers into genomics are discovering again the need for systems of nomenclature-names are what we use ...to communicate about organisms, and by extension the rest of their biology. Here, we briefly outline the history of the published international codes of nomenclature, tracing them from the time of Linnaeus in the eighteenth century to the present day. We then outline some of what we feel are the major challenges that face the codes in the twenty-first century; focusing primarily on publication, priority, typification and the role of science in the naming of organisms. We conclude that the codes are essential for taxonomists in the pursuance of their science, and that the democratic nature of decision-making in the regulation of the rules of nomenclature, though sometimes perceived as a potential weakness, is in fact one of its great strengths.
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