X-ray polarimetry based on gas pixel detectors (GPDs) has reached a high level of maturity thanks to the Imaging X-ray Polarimeter Explorer (IXPE) providing the first-ever spatially resolved ...polarimetric measurements. However, as this a new technique, a few unexpected effects have emerged in the course of in-flight operations. In particular, it was almost immediately found that, the unpolarized calibration sources on-board were showing radially polarized halos. The origin of these features was recognized in a correlation between the error in reconstructing the absorption point of the X-ray photon and the direction of its electric field vector. Here, we present and discuss this effect in detail, showing that it is possible to provide a simple and robust mathematical formalism to handle it. We further show its role and relevance for the recent IXPE measures as well as for the use of GPD-based techniques in general. We also illustrate how to model it in the context of studying extended sources.
Identifying the sources of variation in mating interactions between males and females is important because this variation influences the strength and/or the direction of sexual selection that ...populations experience. While the origins and effects of variation in male attractiveness and ornamentation have received much scrutiny, the causes and consequences of intraspecific variation in females have been relatively overlooked. We used cytogenetic cloning techniques developed for Drosophila melanogaster to create "hemiclonal" males and females with whom we directly observed sexual interaction between individuals of different known genetic backgrounds and measured subsequent reproductive outcomes. Using this approach, we were able to quantify the genetic contribution of each mate to the observed phenotypic variation in biologically important traits including mating speed, copulation duration, and subsequent offspring production, as well as measure the magnitude and direction of intersexual genetic correlation between female choosiness and male attractiveness.
We found significant additive genetic variation contributing to mating speed that can be attributed to male genetic identity, female genetic identity, but not their interaction. Furthermore we found that phenotypic variation in copulation duration had a significant male-associated genetic component. Female genetic identity and the interaction between male and female genetic identity accounted for a substantial amount of the observed phenotypic variation in egg size. Although previous research predicts a trade-off between egg size and fecundity, this was not evident in our results. We found a strong negative genetic correlation between female choosiness and male attractiveness, a result that suggests a potentially important role for sexually antagonistic alleles in sexual selection processes in our population.
These results further our understanding of sexual selection because they identify that genetic identity plays a significant role in phenotypic variation in female behaviour and fecundity. This variation may be potentially due to ongoing sexual conflict found between the sexes for interacting phenotypes. Our unexpected observation of a negative correlation between female choosiness and male attractiveness highlights the need for more explicit theoretical models of genetic covariance to investigate the coevolution of female choosiness and male attractiveness.
In many species, males have the capacity to directly influence (either positively or negatively) the fitness of their mates and offspring, not only via parental care contributions and/or ...precopulatory resource provisioning, but also via the post‐copulatory activity of those substances passed on to their mates in their ejaculates. Here, we examine how an individual male's identity may be related to phenotypic variation in short‐term female fecundity in the model species, Drosophila melanogaster. The effect of male identity on short‐term fecundity stimulation of females was repeatable across time and accounted for over a fifth of the total observed phenotypic variation in fecundity in two independent populations. The functional explanations for these results and the implications for our understanding of the factors that contribute to the adaptive significance of mating preferences and/or sexual conflict are discussed.
Maternal influenza immunisation can reduce morbidity and mortality associated with influenza infection in pregnant women and young infants. We aimed to determine the vaccine efficacy of maternal ...influenza immunisation against maternal and infant PCR-confirmed influenza, duration of protection, and the effect of gestational age at vaccination on vaccine efficacy, birth outcomes, and infant growth up to 6 months of age.
We did a pooled analysis of three randomised controlled trials done in Nepal (2011–2014), Mali (2011–2014), and South Africa (2011–2013). Pregnant women, gestational age 17–34 weeks in Nepal, 28 weeks or more in Mali, and 20–36 weeks in South Africa, were enrolled. Women were randomly assigned 1:1 to a study group, in which they received trivalent inactivated influenza vaccine (IIV) in all three trials, or a control group, in which they received saline placebo in Nepal and South Africa or quadrivalent meningococcal conjugate vaccine in Mali. Enrolment at all sites was complete by April 24, 2013. Infants and women were assessed for respiratory illness, and samples from those that met the case definition were tested for influenza by PCR testing. Growth measurements, including length and weight, were obtained at birth at all sites, at 24 weeks in South Africa, and at 6 months in Nepal and Mali. The three trials are registered with ClinicalTrials.gov, numbers NCT01430689, NCT01034254, and NCT02465190.
10 002 women and 9800 liveborn infants were included. Pooled efficacy of maternal vaccination to prevent infant PCR-confirmed influenza up to 6 months of age was 35% (95% CI 19 to 47). The pooled estimate was 56% (28 to 73) within the first 2 months of life, 39% (11 to 58) between 2 and 4 months, and 19% (–9 to 40) between 4 and 6 months. In women, from enrolment during pregnancy to the end of follow-up at 6 months postpartum, the vaccine was 50% (95% CI 32–63) efficacious against PCR-confirmed influenza. Efficacy was 42% (12 to 61) during pregnancy and 60% (36 to 75) postpartum. In women vaccinated before 29 weeks gestational age, the estimated efficacy was 30% (–2 to 52), and in women vaccinated at or after 29 weeks, efficacy was 71% (50 to 83). Efficacy was similar in infants born to mothers vaccinated before or after 29 weeks gestation (34% 95% CI 12 to 51 vs 35% 11 to 52). There was no overall association between maternal vaccination and low birthweight, stillbirth, preterm birth, and small for gestational age. At 6 months of age, the intervention and control groups were similar in terms of underweight (weight-for-age), stunted (length-for-age), and wasted (weight-for-length). Median centile change from birth to 6 months of age was similar between the intervention and the control groups for both weight and length.
The assessment of efficacy for women vaccinated before 29 weeks gestational age might have been underpowered, because the point estimate suggests that there might be efficacy despite wide CIs. Estimates of efficacy against PCR-confirmed influenza and safety in terms of adverse birth outcomes should be incorporated into any further consideration of maternal influenza immunisation recommendations.
Bill & Melinda Gates Foundation.
We have completed part of a program to study the X-ray emission properties of old rotation-powered pulsars with XMM-Newton in order to probe and identify the origin of their X-radiation. The X-ray ...emission from these old pulsars is largely dominated by nonthermal processes. None of the observed spectra required adding a thermal component consisting of either a hot polar cap or surface cooling emission to model the data. The X-ray spectrum of PSR B0950+08 is best described by a single power law of photon index a = 1.93 super(+) sub(-) super(0) sub(0) super(.) sub(.) super(1) sub(1) super(4) sub(2). Taking optical data from the VLT FORS1 into account, a broken power-law model with the break point E sub(break) = 0.67 super(+) sub(-) super(0) sub(0) super(.) sub(.) super(1) sub(4) super(8) sub(1) keV and the photon indices a sub(1) = 1.27 super(+) sub(-) super(0) sub(0) super(.) sub(.) super(0) sub(0) super(2) sub(1) and a sub(2) = 1.88 super(+) sub(-) super(0) sub(0) super(.) sub(.) super(1) sub(1) super(4) sub(1) for E < E sub(break) and E > E sub(break), respectively, is found to describe the pulsar's broadband spectrum from the optical to the X-ray band. Three s temperature upper limits for possible contributions from a heated polar cap or the whole neutron star surface are T super( ) sub(pc) < 0.87 x 10 super(6) and T super( ) sub(8) < 0.48 x 10 super(6) K, respectively. We also find that the X-ray emission from PSR B0950+08 is pulsed with two peaks per rotation period. The phase separation between the two X-ray peaks is 6144 (maximum to maximum), which is similar to the pulse peak separation observed in the radio band at 1.4 GHz. The main radio peak and the trailing X-ray peak are almost phase-aligned. The fraction of X-ray pulsed photons is 630%. A phase-resolved spectral analysis confirms the nonthermal nature of the pulsed emission and finds no spectral variations as a function of pulse phase. Detailed pulse profile simulations using the polar gap, the outer gap, and the two-pole caustic model constrain the pulsar's emission geometry to be that of an almost orthogonal rotator, for which the two-pole caustic model can reproduce the observed doubly peaked X-ray pulse profile. The spectral emission properties observed for PSR B0823+26 are similar to those of PSR B0950+08. Its energy spectrum is very well described by a single power law with photon index a = 2.5 super(+) sub(-) super(0) sub(0) super(.) sub(.) super(9) sub(45). Three s temperature upper limits for thermal contributions from a hot polar cap or from the entire neutron star surface are T super( ) sub(pc) < 1.17 x 10 super(6) and T super( ) sub(8) < 0.5 x 10 super(6) K, respectively. There is evidence for pulsed X-ray emission at the 697% confidence level with a pulsed fraction of 49% c 22%. For PSR J2043+2740, which is located 61 outside the boundary of the Cygnus Loop, we report the first detection of X-ray emission. A power-law spectrum or a combination of a thermal and a power-law spectrum both yield acceptable descriptions of its X-ray spectrum. No X-ray pulses are detected from PSR J2043+2740 and the sensitivity is low--the 2 s pulsed fraction upper limit is 57% assuming a sinusoidal pulse profile.
We present an X-ray spectropolarimetric analysis of the bright Seyfert galaxy NGC 4151. The source has been observed with the Imaging X-ray Polarimetry Explorer (IXPE) for 700 ks, complemented with ...simultaneous XMM–Newton (50 ks) and NuSTAR (100 ks) pointings. A polarization degree Π = 4.9 ± 1.1 per cent and angle Ψ = 86° ± 7° east of north (68 per cent confidence level) are measured in the 2–8 keV energy range. The spectropolarimetric analysis shows that the polarization could be entirely due to reflection. Given the low reflection flux in the IXPE band, this requires, however, a reflection with a very large (>38 per cent) polarization degree. Assuming more reasonable values, a polarization degree of the hot corona ranging from ∼4 to ∼8 per cent is found. The observed polarization degree excludes a ‘spherical’ lamppost geometry for the corona, suggesting instead a slab-like geometry, possibly a wedge, as determined via Monte Carlo simulations. This is further confirmed by the X-ray polarization angle, which coincides with the direction of the extended radio emission in this source, supposed to match the disc axis. NGC 4151 is the first active galactic nucleus with an X-ray polarization measure for the corona, illustrating the capabilities of X-ray polarimetry and IXPE in unveiling its geometry.
ABSTRACT
We report spectro-polarimetric results of an observational campaign of the bright neutron star low-mass X-ray binary Cyg X-2 simultaneously observed by IXPE, NICER, and INTEGRAL. ...Consistently with previous results, the broad-band spectrum is characterized by a lower-energy component, attributed to the accretion disc with kTin ≈ 1 keV, plus unsaturated Comptonization in thermal plasma with temperature kTe = 3 keV and optical depth τ ≈ 4, assuming a slab geometry. We measure the polarization degree in the 2–8 keV band P = 1.8 ± 0.3 per cent and polarization angle ϕ = 140° ± 4°, consistent with the previous X-ray polarimetric measurements by OSO-8 as well as with the direction of the radio jet which was earlier observed from the source. While polarization of the disc spectral component is poorly constrained with the IXPE data, the Comptonized emission has a polarization degree P = 4.0 ± 0.7 per cent and a polarization angle aligned with the radio jet. Our results strongly favour a spreading layer at the neutron star surface as the main source of the polarization signal. However, we cannot exclude a significant contribution from reflection off the accretion disc, as indicated by the presence of the iron fluorescence line.
ABSTRACT
We report on the Imaging X-ray Polarimetry Explorer (IXPE) observation of the closest and X-ray brightest Compton-thick active galactic nucleus (AGN), the Circinus galaxy. We find the source ...to be significantly polarized in the 2–6 keV band. From previous studies, the X-ray spectrum is known to be dominated by reflection components, both neutral (torus) and ionized (ionization cones). Our analysis indicates that the polarization degree is 28 ± 7 per cent (at 68 per cent confidence level) for the neutral reflector, with a polarization angle of 18° ± 5°, roughly perpendicular to the radio jet. The polarization of the ionized reflection is unconstrained. A comparison with Monte Carlo simulations of the polarization expected from the torus shows that the neutral reflector is consistent with being an equatorial torus with a half-opening angle of 45°–55°. This is the first X-ray polarization detection in a Seyfert galaxy, demonstrating the power of X-ray polarimetry in probing the geometry of the circumnuclear regions of AGNs, and confirming the basic predictions of standard Unification Models.
We report on a comprehensive analysis of simultaneous X-ray polarimetric and spectral data of the bright atoll source GX 9+9 with the Imaging X-ray Polarimetry Explorer (IXPE) and
NuSTAR
. The source ...is significantly polarized in the 4–8 keV band, with a degree of 2.2% ± 0.5% (uncertainty at the 68% confidence level). The
NuSTAR
broad-band spectrum clearly shows an iron line, and is well described by a model including thermal disc emission, a Comptonized component, and reflection. From a spectro-polarimetric fit, we obtain an upper limit to the polarization degree of the disc of 4% (at the 99% confidence level), while the contribution of Comptonized and reflected radiation cannot be conclusively separated. However, the polarization is consistent with resulting from a combination of Comptonization in a boundary or spreading layer, plus reflection off the disc, which significantly contributes in any realistic scenario.
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