In which b(z) is parasite transmissibility, z is parasite virulence, dp is the rate at which parasite offspring 'disperse' to new, randomly chosen patches, i is the number of uninfected hosts, j is the number of infected hosts, class (k, l) is a patch with k uninfected hosts and l infected hosts,r(k,l) is the relatedness between two different parasites on the same class-(k, l) patch, v(k, l) is the reproductive value of a class- (k, l) parasite, and prime denotes differentiation. ... the reduction in optimal virulence does not seem to be the simple "individual-level adaptation" that the authors claim1. ... they suggest that models of meiotic drive could also be used to define individual-level adaptation out of existence2.